Scarabaeidae
The family Scarabaeidae, as currently defined, consists of over 35,000 species of beetles worldwide; they are often called scarabs or scarab beetles. The classification of this family has undergone significant change. Several groups formerly treated as subfamilies have been elevated to family rank, and some reduced to lower ranks. The subfamilies listed in this article are in accordance with those in Catalogue of Life (2023).

Edmund Reitter · Public Domain · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- scarab beetlePBDB
- Sits inside
- ScarabaeoideaPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 121 million years on record.
The record covers at least 121.4 Mya to 11.7 ka. No occurrence read is dated outside 125.77 Mya to 0 Mya.
467 occurrences of Scarabaeidae on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
467 occurrences- Early Cretaceous7%
- Late Cretaceousunder 1%
- Paleoceneunder 1%
- Eocene19%
- Oligocene4%
- Miocene17%
- Pliocene3%
- Pleistocene49%
- Holoceneunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
48 formations
- Highbury Silts and Sands Formation52
- Florissant Formation46
- Shanwang Formation32
- Upper Freshwater-Molasse Formation26
- Zaza Formation16
- Eifel Formation13
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The named rock units Scarabaeidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 24 further formations hold fewer occurrences each and are not listed. 152 occurrences have no formation on record, which is common for older literature.
Scarabaeidae and its relatives
PleistoceneWhere Scarabaeidae and its relatives were recorded in the Pleistocene.
Found in the same rock
6,542 taxa- Aphodius67 sites
- Stenus37 sites
- Aleocharinae32 sites
- Carabidae28 sites
- Bembidion25 sites
- Elateridae25 sites
- Coleoptera24 sites
- Phyllobius24 sites
- Staphylinidae22 sites
- Helophorus21 sites
- Morychus viridis21 sites
- Hydraena20 sites
- Chrysomelidae19 sites
- Curtonotus alpinus19 sites
- Curculionidae18 sites
- Donacia18 sites
- Onthophagus17 sites
- Pterostichus (Cryobius)17 sites
- Apion16 sites
- Dytiscidae16 sites
- Gyrinus16 sites
- Oulimnius troglodytes16 sites
- Poecilus nearcticus16 sites
- Bledius15 sites
The Paleobiology Database records 224 sites for Scarabaeidae, and this reads all of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
124 references- Marine Isotope Stage 9 environments of fluvial deposits at Hackney, north London, UK52
- Late Cenozoic insects of northern Eurasia28
- Zhang, 198921
- New Miocene Coleoptera from Florissant18
- Beiträge zur Insektenfauna Oeningens: Coleoptera. Geodephagen, Hydrocanthariden, Gyriniden, Brachelytren, Clavicornen, Lamellicornen und Buprestiden16
- Systematik, Phylogenie, Taphonomie und Paläoökologie der Insekten aus dem Mittel-Eozän des Eckfelder Maares, Vulkaneifel13
- Middle Pleistocene deposits at Frog Hall Pit, Stretton-on-Dunsmore, Warwickshire, English Midlands, and their implications for the age of the type Wolstonian12
- Heer, 184712
- Late Pleistocene insects from the Mizozono Formation in Yoshimatsu-cho, Kagoshima Prefecture, Japan11
- Palaeoenvironmental interpretation of Middle and Late Wisconsinan fossil coleopteran assemblages from western Olympic Peninsula, Washington, USA11
- The paleoenvironment of the Olympia beds based on fossil beetles from Discovery Park, Seattle, Washington, U.S.A.10
- Environmental interpretation of fossil insect assemblages from MIS 5 at Ziegler Reservoir, Snowmass Village, Colorado10
Each of the 467 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 112 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Scarabaeidae
- Source
- Wikipedia/Commons
- Credit
- Edmund Reitter
- Licence
- Public Domain
- Commercial use
- the licence allows it
No 3D scan of Scarabaeidae was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itScarabaeoidea through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Latreille 1802