Bembidion
Bembidion is the largest genus of beetles in the family Carabidae by number of species. All species are small and move very fast. Most of them live close to water. The genus has a biantitropical distribution, meaning they are found in both the Northern and Southern Hemispheres, but not in the tropics. In warmer regions it is substituted by closely related Tachys and other genera.

Siga · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- BembidiiniPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 33 million years on record.
The record covers at least 33.9 Mya to 11.7 ka. No occurrence read is dated outside 37.71 Mya to 0 Mya.
216 occurrences of Bembidion on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
216 occurrences- Eocene1%
- Miocene6%
- Pleistocene91%
- Holoceneunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
14 formations
- Kap København Formation62
- Highbury Silts and Sands Formation40
- Titusville Till Formation12
- Olympia beds Formation8
- Edoma Formation5
- Imuruk Volcanics Formation3
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The named rock units Bembidion has been recovered from, largest first. Drawn where the rock is now, not where it formed. 72 occurrences have no formation on record, which is common for older literature.
Bembidion and its relatives
PleistoceneWhere Bembidion and its relatives were recorded in the Pleistocene.
Found in the same rock
1,489 taxa- Stenus40 sites
- Aphodius38 sites
- Aleocharinae34 sites
- Pterostichus (Cryobius)27 sites
- Helophorus21 sites
- Phyllobius19 sites
- Bledius18 sites
- Carabidae18 sites
- Chrysomelidae17 sites
- Morychus viridis17 sites
- Dyschirius16 sites
- Elateridae16 sites
- Agabus15 sites
- Curtonotus alpinus15 sites
- Apion14 sites
- Hydroporus14 sites
- Tournotaris bimaculatus14 sites
- Hydraena13 sites
- Hydrobius fuscipes13 sites
- Olophrum consimile13 sites
- Pterostichus13 sites
- Staphylinidae13 sites
- Dyschiriodes globosus12 sites
- Gyrinus12 sites
The Paleobiology Database records 91 sites for Bembidion, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
27 references- Palaeoentomology of the Kap København Formation, a Plio-Pleistocene sequence in Peary Land, North Greenland62
- Marine Isotope Stage 9 environments of fluvial deposits at Hackney, north London, UK40
- Late Cenozoic insects of northern Eurasia25
- Fossil Beetle Evidence for a Short Warm Interval near 40,000 yr B.P. at Titusville, Pennsylvania12
- Palaeoenvironmental interpretation of Middle and Late Wisconsinan fossil coleopteran assemblages from western Olympic Peninsula, Washington, USA9
- A Late Pleistocene fauna and flora from Upton Warren, Worcestershire9
- The paleoenvironment of the Olympia beds based on fossil beetles from Discovery Park, Seattle, Washington, U.S.A.8
- Environmental interpretation of fossil insect assemblages from MIS 5 at Ziegler Reservoir, Snowmass Village, Colorado8
- Middle Pleistocene deposits at Frog Hall Pit, Stretton-on-Dunsmore, Warwickshire, English Midlands, and their implications for the age of the type Wolstonian6
- Late Devensian river deposits beneath the floodplain terrace of the River Thames at Abingdon, Berkshire, England5
- A diverse late-glacial (Mackinaw Phase) biota from Leamington, Ontario4
- Paleogeography of the lower Volga at the end of Atelian time based on entomological and botanical data3
Each of the 216 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 15 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content2
Picture of Bembidion
- Source
- Wikipedia/Commons
- Credit
- Siga
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
Photogrammetry
- Source
- MorphoSource
- Licence
- not stated by the source
- Commercial use
- cannot be read from the licence given — check the record
- Specimen
- Bembidion Bembidion compactum · ETHZ:ENT:0577233
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Bembidion”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itBembidiini through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Latreille 1802