Stenus
Stenus is a genus of semiaquatic rove beetles in the subfamily Steninae, and one of the largest genera in the kingdom Animalia, with some 3100 known species worldwide. They are predators of Collembola and other small arthropods. Adults have a protrusible labium with a sticky tip used in prey capture. To overcome the rapid escape of Collembola, the labium is protruded at high speed by hemolymph pressure, and immediately withdrawn, pulling the prey within the range of the mandibles. However, the labium tip does not easily stick to prey covered in scales or setae or that have a large body size. Stenus comma is more likely to catch such prey by lunging forward and grabbing them directly with its mandibles rather than using its labium. Stenus species are also known for "skimming" on the water surface using their pygidial gland secretions that act as a surfactant and rapidly propel the beetle fast forward, a phenomenon known as the Marangoni effect. Stenus comma has been seen to achieve a velocity of 0.75 m/s, and to cover a distance of up to 15 m if the secretion is continuous.

Udo Schmidt from Deutschland · CC BY-SA 2.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- SteninaePBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 93 million years on record.
The record covers at least 93.9 Mya to 129 ka. No occurrence read is dated outside 100.5 Mya to 11.7 ka.
110 occurrences of Stenus on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
110 occurrences- Late Cretaceous2%
- Eocene20%
- Oligocene5%
- Miocene2%
- Pleistocene72%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
11 formations
- Kap København Formation23
- Highbury Silts and Sands Formation12
- Titusville Till Formation6
- Florissant Formation4
- Quadra Sand Formation3
- Niveau du gypse d'Aix Formation2
Loading coastlines…
The named rock units Stenus has been recovered from, largest first. Drawn where the rock is now, not where it formed. 54 occurrences have no formation on record, which is common for older literature.
Stenus and its relatives
PleistoceneWhere Stenus and its relatives were recorded in the Pleistocene.
Found in the same rock
2,843 taxa- Aleocharinae42 sites
- Aphodius31 sites
- Helophorus29 sites
- Bembidion23 sites
- Bledius18 sites
- Hydraena17 sites
- Apion15 sites
- Arpedium cribratum15 sites
- Elateridae15 sites
- Scydmaeninae15 sites
- Agathidium14 sites
- Dytiscidae14 sites
- Dyschirius13 sites
- Oulimnius troglodytes13 sites
- Oulimnius tuberculatus13 sites
- Oxytelus laqueatus13 sites
- Rhynchaenus quercus13 sites
- Anotylus rugosus12 sites
- Carabidae12 sites
- Chrysomelidae12 sites
- Dyschiriodes globosus12 sites
- Gyrinus12 sites
- Bryaxis11 sites
- Carpelimus11 sites
The Paleobiology Database records 76 sites for Stenus, and this reads all of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
24 references- Palaeoentomology of the Kap København Formation, a Plio-Pleistocene sequence in Peary Land, North Greenland23
- Stenus Latreille, 1797 aus dem Baltischen Bernstein nebst Bemerkungen über andere fossile Stenus-Arten (Coleoptera, Staphylinidae)17
- Palaeoenvironmental interpretation of Middle and Late Wisconsinan fossil coleopteran assemblages from western Olympic Peninsula, Washington, USA16
- Marine Isotope Stage 9 environments of fluvial deposits at Hackney, north London, UK12
- Middle Pleistocene deposits at Frog Hall Pit, Stretton-on-Dunsmore, Warwickshire, English Midlands, and their implications for the age of the type Wolstonian8
- Fossil Beetle Evidence for a Short Warm Interval near 40,000 yr B.P. at Titusville, Pennsylvania6
- Environmental interpretation of fossil insect assemblages from MIS 5 at Ziegler Reservoir, Snowmass Village, Colorado3
- Late Cenozoic insects of northern Eurasia3
- Pre-Vashon fossil Coleoptera of Fraser age from the Fraser Lowland, British Columbia3
- Late Devensian river deposits beneath the floodplain terrace of the River Thames at Abingdon, Berkshire, England3
- The paleoenvironment of the Olympia beds based on fossil beetles from Discovery Park, Seattle, Washington, U.S.A.2
- Adephagous and clavicorn Coleoptera from the Tertiary deposits at Florissant, Colorado with descriptions of a few other forms and a systematic list of the non-rhynchophorus Tertiary Coleoptera of North America2
Each of the 110 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 12 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content2
Picture of Stenus
- Source
- Wikipedia/Commons
- Credit
- Udo Schmidt from Deutschland
- Licence
- CC BY-SA 2.0
- Commercial use
- the licence allows it
Photogrammetry
- Source
- MorphoSource
- Licence
- not stated by the source
- Commercial use
- cannot be read from the licence given — check the record
- Specimen
- Stenus Stenus guttula · ETHZ:ENT:0576222
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Stenus”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itSteninae through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Latreille 1796