Scydmaeninae
Scydmaeninae are a subfamily of small beetles, commonly called ant-like stone beetles or scydmaenines. These beetles occur worldwide, and the subfamily includes some 4,500 species in about 80 genera. Established as a family, they were reduced in status to a subfamily of Staphylinidae in 2009

IES MANUEL GARCÍA BARROS A ESTRADA- PONTEVEDRA · CC BY-SA 2.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- antlike stone beetlePBDB
- Sits inside
- StaphylinidaePBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 109 million years on record.
The record covers at least 110.1 Mya to 129 ka. No occurrence read is dated outside 113.2 Mya to 11.7 ka.
143 occurrences of Scydmaeninae on record, most of them in the Eocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
143 occurrences- Early Cretaceous3%
- Late Cretaceous36%
- Eocene41%
- Oligocene1%
- Miocene6%
- Plioceneunder 1%
- Pleistocene12%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
7 formations
- Titusville Till Formation4
- Las Penosas Formation2
- Kheta Formation2
- Foremost Formation · Belly River Group1
- Imuruk Volcanics Formation1
- Niveau du gypse d'Aix Formation1
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The named rock units Scydmaeninae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 131 occurrences have no formation on record, which is common for older literature.
Scydmaeninae and its relatives
EoceneWhere Scydmaeninae and its relatives were recorded in the Eocene.
Found in the same rock
4,069 taxa- Stenus18 sites
- Aleocharinae15 sites
- Helophorus12 sites
- Sthereus quadrituberculatus11 sites
- Agathidium10 sites
- Arpedium cribratum10 sites
- Dytiscidae10 sites
- Elateridae9 sites
- Oropus9 sites
- Ptiliidae9 sites
- Bembidion8 sites
- Pseudohylesinus nebulosus8 sites
- Trechus (Trechus) oregonensis8 sites
- Aphodius7 sites
- Euaesthetinae7 sites
- Pelecomalium testaceum7 sites
- Phloeotribus lecontei7 sites
- Pityophthorus nitidulus7 sites
- Sonoma7 sites
- Staphylinidae7 sites
- Staphylininae7 sites
- Gymnusa6 sites
- Hydrophilidae6 sites
- Olophrum boreale6 sites
The Paleobiology Database records 61 sites for Scydmaeninae, and this reads all of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
57 references- Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber25
- Palaeoenvironmental interpretation of Middle and Late Wisconsinan fossil coleopteran assemblages from western Olympic Peninsula, Washington, USA11
- Die Scydmaeniden des baltischen Bernsteines11
- New ant-like stone beetles in mid-Cretaceous amber from Myanmar (Coleoptera: Staphylinidae: Scydmaeninae)6
- Evolution of Mastigitae: Mesozoic and Cenozoic fossils crucial for reclassification of extant tribes (Coleoptera: Staphylinidae: Scydmaeninae)5
- Discrimination of species of Hapsomela Poinar & Brown (Coleoptera: Staphylinidae: Scydmaeninae) in mid-Cretaceous Burmese amber4
- Fossil Beetle Evidence for a Short Warm Interval near 40,000 yr B.P. at Titusville, Pennsylvania4
- Fossile Scydmaenidae in Baltischem und Dominikanischem Bernstein (Coleoptera: Scydmaenidae)4
- Notes on mid-Cretaceous †Pangusyndicus and its relatives, with description of †Loeblitoides gen. nov. and a †Nuegua-like new species (Coleoptera: Staphylinidae: Scydmaeninae)3
- The Mastigitae genus †Baltostigus in Upper Eocene Rovno amber (Coleoptera: Staphylinidae: Scydmaeninae)3
- †Praphennium gen. nov.: Extant mite killers meet their Cenomanian relatives (Coleoptera: Staphylinidae: Scydmaeninae: Cephenniini)3
- The genus Scydmobisetia Jałoszyński & Yamamoto, 2016 in Upper Cretaceous Burmese amber (Coleoptera, Staphylinidae, Scydmaeninae)3
Each of the 143 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 45 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Scydmaeninae
- Source
- Wikipedia/Commons
- Credit
- IES MANUEL GARCÍA BARROS A ESTRADA- PONTEVEDRA
- Licence
- CC BY-SA 2.0
- Commercial use
- the licence allows it
No 3D scan was found. Smithsonian Open Access, MorphoSource and Sketchfab were all searched.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itStaphylinidae through timeFormations of the EoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Leach 1815)