Coleoptera
Beetles are insects that form the order Coleoptera, in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 described species, is the largest of all orders, constituting almost 40% of described arthropods and 25% of all known animal species; new species are discovered frequently, with estimates suggesting that there are between 0.9 and 2.1 million total species.

Bugboy52.40 · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- beetlePBDB
- Sits inside
- ColeopteridaPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 293 million years on record.
The record covers at least 293.52 Mya to 11.7 ka. No occurrence read is dated outside 298.9 Mya to 0 Mya.
12,000+ occurrences of Coleoptera on record. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
12,000 occurrences read- Permian2%
- Early Triassicunder 1%
- Middle Triassic1%
- Late Triassic2%
- Early Jurassic3%
- Middle Jurassic1%
- Late Jurassic7%
- Early Cretaceous9%
- Late Cretaceous1%
- Paleocene1%
- Eocene20%
- Oligocene4%
- Miocene9%
- Pliocene2%
- Pleistocene37%
- Holoceneunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. The read hit its cap, so every figure above is a floor and the shares are of the occurrences read. More on what this infers.
Found in
at least 352 formations
- Florissant Formation821
- Highbury Silts and Sands Formation735
- Karabastau Formation542
- Kap København Formation469
- Titusville Till Formation312
- Upper Freshwater-Molasse Formation267
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The named rock units Coleoptera has been recovered from, largest first. Drawn where the rock is now, not where it formed. The read hit its cap, so each count is a floor. 328 further formations hold fewer occurrences each and are not listed. 3,897 occurrences have no formation on record, which is common for older literature.
Found in the same rock
7,104 taxa- Blattodea11 sites
- Curculionidae11 sites
- Odonata9 sites
- Diptera8 sites
- Gastropoda8 sites
- Heteroptera8 sites
- Protoscelis jurassica8 sites
- Archirhyphus asiaticus7 sites
- Equisetum7 sites
- Liassothrips crassipes7 sites
- Protoscelis longicornis7 sites
- Taeniopteris7 sites
- Apioninae6 sites
- Camponotites lignitus6 sites
- Carabidae6 sites
- Curculionoidea6 sites
- Distenorrhinus (Distenorrhinus) pallidirostris6 sites
- Elateridae6 sites
- Hydrophilidae6 sites
- Insecta6 sites
- Kararhynchus occiduus6 sites
- Osteichthyes6 sites
- Panorpidium geinitzi6 sites
- Paramesopsocus adibi6 sites
The Paleobiology Database records at least 490 sites for Coleoptera, and this reads 240 of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
at least 1,226 references- Late Cenozoic insects of northern Eurasia731
- Marine Isotope Stage 9 environments of fluvial deposits at Hackney, north London, UK722
- Palaeoentomology of the Kap København Formation, a Plio-Pleistocene sequence in Peary Land, North Greenland469
- Palaeoenvironmental interpretation of Middle and Late Wisconsinan fossil coleopteran assemblages from western Olympic Peninsula, Washington, USA430
- Fossil Beetle Evidence for a Short Warm Interval near 40,000 yr B.P. at Titusville, Pennsylvania310
- Tertiary Rhynchophorous Coleoptera of the United States228
- Middle Pleistocene deposits at Frog Hall Pit, Stretton-on-Dunsmore, Warwickshire, English Midlands, and their implications for the age of the type Wolstonian205
- A Late Pleistocene fauna and flora from Upton Warren, Worcestershire181
- The paleoenvironment of the Olympia beds based on fossil beetles from Discovery Park, Seattle, Washington, U.S.A.180
- Mezozoiskie zhestkokryiye164
- Fossil click-beetles (Coleoptera, Elateridae) from the Upper Jurassic of Karatau146
- Systematik, Phylogenie, Taphonomie und Paläoökologie der Insekten aus dem Mittel-Eozän des Eckfelder Maares, Vulkaneifel142
Each of the 12,000 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. That read hit its cap, so each count is a floor. 1,214 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content2
Picture of Coleoptera
- Source
- Wikipedia/Commons
- Credit
- Bugboy52.40
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
3D model
- Source
- Sketchfab
- Licence
- CC Attribution-NonCommercial
- Commercial use
- the licence does not allow it
The description on this site
- Source
- Wikipedia, “Beetle”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itColeopterida through timeThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Linnaeus 1758