Scarabaeinae
The scarab beetle subfamily Scarabaeinae consists of species collectively called true dung beetles. Most of the beetles of this subfamily feed exclusively on dung. However, some may feed on decomposing matter including carrion, decaying fruits and fungi. Dung beetles can be placed into three structural guilds based on their method of dung processing namely rollers (telecoprids), dwellers (endocoprids) and tunnelers (paracoprids). Dung removal and burial by dung beetles result in ecological benefits such as soil aeration and fertilization; improved nutrient cycling and uptake by plants, increase in pasture quality, biological control of pest flies and intestinal parasites and secondary seed dispersal. Well-known members include the genera Scarabaeus and Sisyphus, and Phanaeus vindex.

Didier Descouens · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- ScarabaeidaePBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 59 million years on record.
The record covers at least 59.24 Mya to 129 ka. No occurrence read is dated outside 66 Mya to 11.7 ka.
70 occurrences of Scarabaeinae on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
70 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
17 formations
- Highbury Silts and Sands Formation11
- Upper Freshwater-Molasse Formation7
- Mizozono Formation4
- Tsubusagawa Formation3
- Hiwegi Formation2
- Shanwang Formation2
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The named rock units Scarabaeinae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 28 occurrences have no formation on record, which is common for older literature.
Scarabaeinae and its relatives
PleistoceneWhere Scarabaeinae and its relatives were recorded in the Pleistocene.
Found in the same rock
2,014 taxa- Coleoptera18 sites
- Onthophagus18 sites
- Carabidae14 sites
- Aphodius11 sites
- Curculionidae11 sites
- Staphylinidae10 sites
- Chrysomelidae8 sites
- Coelostoma8 sites
- Elateridae8 sites
- Gyrinus8 sites
- Aleocharinae7 sites
- Donaciinae7 sites
- Limnius volckmari7 sites
- Melanotus punctolineatus7 sites
- Scarabaeidae7 sites
- Bryaxis6 sites
- Carpelimus6 sites
- Dyschiriodes globosus6 sites
- Heteroptera6 sites
- Hydraena6 sites
- Oulimnius troglodytes6 sites
- Oulimnius tuberculatus6 sites
- Pelochares versicolor6 sites
- Pomatinus substriatus6 sites
The Paleobiology Database records 45 sites for Scarabaeinae, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
32 references- Marine Isotope Stage 9 environments of fluvial deposits at Hackney, north London, UK11
- Fossil arthropods of California 11. Descriptions of the dung beetles (Scarabaeidae) of the tar pits6
- Beiträge zur Insektenfauna Oeningens: Coleoptera. Geodephagen, Hydrocanthariden, Gyriniden, Brachelytren, Clavicornen, Lamellicornen und Buprestiden5
- Late Pleistocene insects from the Mizozono Formation in Yoshimatsu-cho, Kagoshima Prefecture, Japan4
- Reevaluation of Pleistocene scarab beetles from Rancho La Brea, California (Coleoptera: Scarabaeidae)4
- Fossil insects and their paleoenvironments of the Pliocene Tsubusagawa Formation in Ajimu Basin, Oita Prefecture, Kyushu, Japan3
- The Clark's Cave bone deposit and the Late Pleistocene paleoecology of the central Appalachian Mountains of Virginia3
- Three fossil dung beetles (Coleoptera: Scarabaeidae) from the Kenya Miocene3
- A new extinct species of Scybalophagus dung beetle supports the collateral extinction hypothesis at the Chilean South American Pleistocene-Holocene boundary2
- A review and phylogeny of Scarabaeine dung beetle fossils (Coleoptera: Scarabaeidae: Scarabaeinae), with the description of two Canthochilum species from Dominican amber2
- Systematische und palökologische Untersuchungen an Insekten aus dem Mittel-Eozän der Grube messel bei Darmstadt2
- Zhang, 19892
Each of the 70 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 20 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content2
Picture of Scarabaeinae
- Source
- Wikipedia/Commons
- Credit
- Didier Descouens
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
3D model
- Source
- Sketchfab
- Credit
- Binkley-Spacetrucker
- Licence
- CC Attribution
- Commercial use
- cannot be read from the licence given — check the record
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itScarabaeidae through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Latreille 1802