Natricinae
The Natricinae are a subfamily of colubroid snakes, sometimes referred to as a family (Natricidae). The subfamily comprises 36 genera. Members include many very common snake species, such as the European grass snakes, and the North American water snakes and garter snakes. Some Old World members of the subfamily are known as keelbacks, because their dorsal scales exhibit strong keeling.

Orchi · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- ColubridaePBDB
- Diet
- carnivorePBDB, for Colubridae
- Life habit
- ground dwellingPBDB, for Colubridae
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Amniota
Time the record covers
At least 23 million years on record.
The record covers at least 23.04 Mya to 11.7 ka. No occurrence read is dated outside 27.3 Mya to 0 Mya.
181 occurrences of Natricinae on record, most of them in the Miocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
181 occurrences- Oligoceneunder 1%
- Miocene46%
- Pliocene12%
- Pleistocene40%
- Holocene1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
31 formations
- Ash Hollow Formation8
- Valentine Formation · Ogallala Group7
- Kingsdown Formation6
- McPherson Formation6
- Bermont Formation4
- Fleming Formation4
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The named rock units Natricinae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 7 further formations hold fewer occurrences each and are not listed. 110 occurrences have no formation on record, which is common for older literature.
Natricinae and its relatives
MioceneWhere Natricinae and its relatives were recorded in the Miocene.
Found in the same rock
2,217 taxa- Rana40 sites
- Thamnophis38 sites
- Nerodia34 sites
- Colubrinae33 sites
- Equus26 sites
- Rana pipiens22 sites
- Bufo21 sites
- Nerodia sipedon20 sites
- Vipera20 sites
- Coluber constrictor19 sites
- Elaphe19 sites
- Lampropeltis getulus19 sites
- Ophisaurus19 sites
- Eumeces18 sites
- Lacerta18 sites
- Colubridae17 sites
- Heterodon platyrhinos17 sites
- Odocoileus virginianus17 sites
- Microtus pennsylvanicus16 sites
- Agkistrodon15 sites
- Ambystoma tigrinum15 sites
- Coluber15 sites
- Lacertidae15 sites
- Lepisosteus15 sites
The Paleobiology Database records 130 sites for Natricinae, and this reads all of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
105 references- High-handed opinions on taxonomy and other matters13
- North American Watersnakes: A Natural History7
- Herpetofauna of the Sandahl Local fauna (Pleistocene: Illinoian) of Kansas6
- Fossil snakes of the Clarendonian (late Miocene) Pratt Slide local fauna of Nebraska, with the description of a new natricine colubrid5
- Snakes of the lower/middle Miocene transition at Vieux Collonges (Rhône, France), with comments on the colonisation of western Europe by colubroids.5
- Herpetofaunas of the Big Springs and Hornet's Nest quarries (northeastern Nebraska, Pleistocene: Late Blancan)5
- Pleistocene amphibians and reptiles from the Leisey Shell Pit, Hillsborough County, Florida4
- Early Pliocene fossil snakes (Squamata, Colubroidea) with various teeth from the fissure deposit in Queshan, Henan, China3
- Amphibians and reptiles from the late Miocene and early Pliocene of the Ptolemais area (Western Macedonia, Greece)3
- A Late Pleistocene Hepetofauna from Clark Quarry, Glynn County, Georgia3
- Snake fauna (Reptilia: Serpentes) from the Early/Middle Miocene of Sandelzhausen and Rothenstein 13 (Germany)3
- Amphibians and squamate reptiles from the late early Miocene (MN 4) of Béon 1 (Montréal-du-Gers, southwestern France)3
Each of the 181 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 93 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Natricinae
- Source
- Wikipedia/Commons
- Credit
- Orchi
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
No 3D scan was found. Smithsonian Open Access, MorphoSource and Sketchfab were all searched.
The description on this site
- Source
- Wikipedia, “Natricinae”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itColubridae through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Bonaparte 1838