Juglans
Walnut trees are any species of tree in the plant genus Juglans, the type genus of the family Juglandaceae, the seeds of which are referred to as walnuts. All species are deciduous trees, 10–40 metres (33–131 ft) tall, with pinnate leaves 200–900 millimetres (7.9–35.4 in), with 5–25 leaflets; the shoots have chambered pith, a character shared with the wingnuts (Pterocarya), but not the hickories (Carya) in the same family.

Bruce Marlin · CC BY 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- walnutPBDB
- Sits inside
- JuglandaceaePBDB
- Diet
- "photoautotroph"PBDB, for Angiospermae
Time the record covers
At least 83 million years on record.
The record covers at least 83.6 Mya to 11.7 ka. No occurrence read is dated outside 100.5 Mya to 0 Mya.
142 occurrences of Juglans on record, most of them in the Miocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
142 occurrences- Late Cretaceous4%
- Paleocene6%
- Eocene18%
- Oligocene7%
- Miocene28%
- Pliocene12%
- Pleistocene18%
- Holocene6%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
44 formations
- Sepulcher Formation10
- Fort Union Formation5
- Okhonchevayamskaya Formation4
- Raton Formation3
- Calvert Formation · Chesapeake Group3
- Shanwang Formation3
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The named rock units Juglans has been recovered from, largest first. Drawn where the rock is now, not where it formed. 20 further formations hold fewer occurrences each and are not listed. 69 occurrences have no formation on record, which is common for older literature.
Juglans and its relatives
MioceneWhere Juglans and its relatives were recorded in the Miocene.
Found in the same rock
1,469 taxa- Alnus61 sites
- Quercus56 sites
- Ulmus50 sites
- Betula42 sites
- Acer41 sites
- Picea41 sites
- Pinus40 sites
- Poaceae38 sites
- Salix38 sites
- Corylus36 sites
- Carya35 sites
- Abies31 sites
- Cyperaceae31 sites
- Asteraceae29 sites
- Carpinus29 sites
- Liquidambar27 sites
- Pterocarya26 sites
- Myrica24 sites
- Tsuga24 sites
- Typha24 sites
- Apiaceae22 sites
- Artemisia22 sites
- Ilex22 sites
- Rosaceae22 sites
The Paleobiology Database records 116 sites for Juglans, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
48 references- A Catalogue of Fossil Plants from India Part 6: Cenozoic (Quaternary) Palynology and Palaeobotany22
- A catalogue of Cenozoic megafossil plants in China11
- Fossil Flora of the Yellowstone National Park11
- Niche evolution versus niche conservatism and habitat loss determine persistence and extirpation in late Neogene European Fagaceae8
- Paleocene flora of the Rocky Mountains and Great Plains8
- Raschleneniye oligotsen-nizhnemiotsenovykh tolshch vostochnoy Kamchatki (zaliv Korfa). Subdivision of Oligocene-lower Miocene layers of eastern Kamchatka, Gulf of Korfa6
- Kleinsäugerstratigraphie in der miozänen Hörnlischüttung (Ostschweiz); (Biostratigraphy of small mammals in the Hörnlischüttung of Miocene age (East Switzerland))5
- Fossil macrospores and massulae of Salvinia natans from the Pliocene and the Quaternary sediments in the Kinki and Tokai districts, Japan5
- Geology and biostratigraphy of the Potomac River Cliffs at Stratford Hall, Westmoreland County, Virginia4
- Die mittelmiozaenen und obermiozaenen Floren aus der Meuroer und Raunoer Folge in der Lausitz. III. Fundstellen und Palaeobiologie4
- Late Miocene and Pliocene Floras in Central Honshu, Japan4
- Reconstructing Neogene vegetation and climates to infer tectonic uplift in western Yunnan, China3
Each of the 142 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 36 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content3
Picture of Juglans
- Source
- Wikipedia/Commons
- Credit
- Bruce Marlin
- Licence
- CC BY 3.0
- Commercial use
- the licence allows it
CT scan
- Source
- MorphoSource
- Licence
- In Copyright
- Commercial use
- cannot be read from the licence given — check the record
- Specimen
- Juglans bergomensis · uwbm:36885-B5585
3D model
- Source
- Sketchfab
- Licence
- CC Attribution
- Commercial use
- cannot be read from the licence given — check the record
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Juglans”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itJuglandaceae through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Linnaeus 1753