Carpinus
Hornbeams are hardwood trees in the plant genus Carpinus in the family Betulaceae. Its species occur across much of the temperate regions of the Northern Hemisphere.

MPF · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- hornbeamPBDB
- Sits inside
- BetulaceaePBDB
- Diet
- "photoautotroph"PBDB, for Angiospermae
Time the record covers
At least 55 million years on record.
The record covers at least 56 Mya to 11.7 ka. No occurrence read is dated outside 66 Mya to 0 Mya.
179 occurrences of Carpinus on record, most of them in the Miocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
179 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
32 formations
- Itahana Formation · Tomioka Group10
- Shanwang Formation8
- Ogawa Formation5
- Motojuku Formation4
- Seto Porcelain Clay Formation · Seto Group3
- Akeyo Formation · Mizunami Group3
Loading coastlines…
The named rock units Carpinus has been recovered from, largest first. Drawn where the rock is now, not where it formed. 8 further formations hold fewer occurrences each and are not listed. 112 occurrences have no formation on record, which is common for older literature.
Carpinus and its relatives
MioceneWhere Carpinus and its relatives were recorded in the Miocene.
Found in the same rock
1,137 taxa- Alnus45 sites
- Ulmus41 sites
- Quercus40 sites
- Betula37 sites
- Acer35 sites
- Corylus34 sites
- Salix34 sites
- Poaceae29 sites
- Juglans27 sites
- Picea27 sites
- Abies26 sites
- Pinus24 sites
- Tilia22 sites
- Cyperaceae21 sites
- Fagus21 sites
- Fraxinus21 sites
- Celtis19 sites
- Potamogeton19 sites
- Rhus19 sites
- Artemisia18 sites
- Asteraceae18 sites
- Apiaceae17 sites
- Carya17 sites
- Populus17 sites
The Paleobiology Database records 75 sites for Carpinus, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
41 references- Die mittelmiozaenen und obermiozaenen Floren aus der Meuroer und Raunoer Folge in der Lausitz. III. Fundstellen und Palaeobiologie38
- Late Miocene and Pliocene Floras in Central Honshu, Japan28
- A catalogue of Cenozoic megafossil plants in China18
- A Catalogue of Fossil Plants from India Part 6: Cenozoic (Quaternary) Palynology and Palaeobotany16
- Die Floren des Haselbacher Serie im Weisselster-Becken (Bezirk Leipzig, DDR)12
- La stratigraphie du remplissage de la Grotte du Vallonnet.5
- Tretichnye Flory Zapadnoi Sibiri5
- Allochthonous seed and fruit floras from the Pliocene of the Lower Rhine Basin4
- Niche evolution versus niche conservatism and habitat loss determine persistence and extirpation in late Neogene European Fagaceae3
- Pleistocene fossil leaflets of Albizia kalkora (Roxb.) Prain (Leguminosae subfamily Mimosoideae) from central Honshu, Japan, and its implications for historical biogeography3
- Die pliozaenen Floren von Thueringen, Deutsche Demokratische Republik3
- Fossil macrospores and massulae of Salvinia natans from the Pliocene and the Quaternary sediments in the Kinki and Tokai districts, Japan3
Each of the 179 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 29 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content2
Picture of Carpinus
- Source
- Wikipedia/Commons
- Credit
- MPF
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Hornbeam”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itBetulaceae through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Linnaeus 1753