Ilex
Ilex or holly is a genus of over 570 species of flowering plants in the family Aquifoliaceae, and the only living genus in that family. Ilex has the most species of any woody dioecious angiosperm genus. The species are evergreen or deciduous trees, shrubs, and climbers from tropics to temperate zones worldwide. The type species is Ilex aquifolium, the common European holly used in Christmas decorations and cards.

Jürgen Howaldt · CC BY-SA 2.0 de · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- hollyPBDB
- Sits inside
- AquifoliaceaePBDB
- Diet
- "photoautotroph"PBDB, for Angiospermae
Time the record covers
At least 93 million years on record.
The record covers at least 93.9 Mya to 11.7 ka. No occurrence read is dated outside 100.5 Mya to 0 Mya.
190 occurrences of Ilex on record, most of them in the Miocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
190 occurrences- Late Cretaceous3%
- Paleocene1%
- Eocene4%
- Oligocene4%
- Miocene46%
- Pliocene23%
- Pleistocene17%
- Holocene3%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
28 formations
- Gatun Formation6
- Culebra Formation5
- Itahana Formation · Tomioka Group4
- Calvert Formation · Chesapeake Group4
- Okinoyama Formation · Ube Group3
- Santa Maria di Ciciliano Formation3
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The named rock units Ilex has been recovered from, largest first. Drawn where the rock is now, not where it formed. 4 further formations hold fewer occurrences each and are not listed. 139 occurrences have no formation on record, which is common for older literature.
Ilex and its relatives
MioceneWhere Ilex and its relatives were recorded in the Miocene.
Found in the same rock
1,398 taxa- Poaceae48 sites
- Asteraceae42 sites
- Cyathea35 sites
- Podocarpus34 sites
- Cyperaceae33 sites
- Alnus32 sites
- Myrtaceae32 sites
- Quercus32 sites
- Symplocos30 sites
- Ericaceae29 sites
- Lycopodium28 sites
- Pinus28 sites
- Carya27 sites
- Liquidambar27 sites
- Acacia24 sites
- Apiaceae24 sites
- Betula24 sites
- Myriophyllum24 sites
- Plantago24 sites
- Sphagnum24 sites
- Syzygium24 sites
- Chenopodiaceae23 sites
- Elaeocarpus23 sites
- Juglans23 sites
The Paleobiology Database records 108 sites for Ilex, and this reads all of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
49 references- Die mittelmiozaenen und obermiozaenen Floren aus der Meuroer und Raunoer Folge in der Lausitz. III. Fundstellen und Palaeobiologie29
- Allochthonous seed and fruit floras from the Pliocene of the Lower Rhine Basin16
- Early Pleistocene vegetation change in upland south-eastern Australia14
- Miocene floras in the lower Rhenish basin and their ecological interpretation13
- A Catalogue of Fossil Plants from India Part 6: Cenozoic (Quaternary) Palynology and Palaeobotany11
- Niche evolution versus niche conservatism and habitat loss determine persistence and extirpation in late Neogene European Fagaceae9
- Late Miocene and Pliocene Floras in Central Honshu, Japan9
- Stratigraphic, Paleoclimatic, and Paleobiogeographic Significance of Tertiary Sporomorphs from Massachusetts9
- Studies in Neotropical Paleobotany. X. The Pliocene communities of Panama - Composistion, numerical representations, and paleocommunity paleoenvironmental reconstructions.7
- A complete pollen record of the last 230 ka from Lynch's Crater, north-eastern Australia6
- Studies in Neotropical Paleobotany. V. The Lower Miocene Communities of Panama - The Culebra Formation.5
- Die pliozaenen Floren von Thueringen, Deutsche Demokratische Republik5
Each of the 190 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 37 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content2
Picture of Ilex
- Source
- Wikipedia/Commons
- Credit
- Jürgen Howaldt
- Licence
- CC BY-SA 2.0 de
- Commercial use
- the licence allows it
CT scan
- Source
- MorphoSource
- Licence
- not stated by the source
- Commercial use
- cannot be read from the licence given — check the record
- Specimen
- Sachatamia ilex · KU:KUH:221613
The description on this site
- Source
- Wikipedia, “Holly”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itAquifoliaceae through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Linnaeus 1753