Harpalinae

Harpalinae is the largest subfamily of ground beetles, containing more than 19,000 species worldwide.

Harpalinae

Uncredited · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.

On record

Recorded, not inferred
Sits inside
CarabidaePBDB
Motility
actively mobilePBDB, for Arthropoda
Environment
terrestrialPBDB, for Insecta

Time the record covers

At least 113 million years on record.

The record covers at least 113.2 Mya to 11.7 ka. No occurrence read is dated outside 132.6 Mya to 0 Mya.

Early Cretaceous — about 2 of 548 occurrencesEarly CretaceousLate Cretaceous — about 7 of 548 occurrencesLate CretaceousPaleocene — about 3 of 548 occurrencesEocene — about 63 of 548 occurrencesEoceneOligocene — about 25 of 548 occurrencesMiocene — about 47 of 548 occurrencesMiocenePliocene — about 12 of 548 occurrencesPleistocene — about 386 of 548 occurrences135–131 Mya — about 0.1 occurrences131–126 Mya — about 0.2 occurrences126–122 Mya — about 0.2 occurrences122–117 Mya — about 0.6 occurrences117–113 Mya — about 0.8 occurrences104–99 Mya — about 0.4 occurrences99–95 Mya — about 1 occurrence95–90 Mya — about 5 occurrences90–86 Mya — about 0.4 occurrences68–63 Mya — about 0.4 occurrences63–59 Mya — about 2 occurrences59–54 Mya — about 2 occurrences54–50 Mya — about 2 occurrences50–45 Mya — about 3 occurrences45–41 Mya — about 1 occurrence41–36 Mya — about 20 occurrences36–32 Mya — about 39 occurrences32–27 Mya — about 7 occurrences27–23 Mya — about 14 occurrences23–18 Mya — about 4 occurrences18–14 Mya — about 6 occurrences14–9.0 Mya — about 28 occurrences9.0–4.5 Mya — about 11 occurrences4.5–0 Mya — about 399 occurrences113.2 Mya11.7 ka135 Mya0 Mya

548 occurrences of Harpalinae on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.

When

548 occurrences

These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.

Found in

39 formations

Kap København — 82.47°N, 23.66°WKap København — 82.45°N, 24.10°WKap København — 82.32°N, 23.85°WKap København — 82.35°N, 24.05°WKap København — 82.46°N, 23.25°WKeremesit — 69.50°N, 158.50°EKeremesit — 69.17°N, 158.12°EHighbury Silts and Sands — 51.56°N, 0.06°WFlorissant — 38.92°N, 105.28°WFlorissant — 38.93°N, 105.24°WEdoma — 69.50°N, 158.50°EUpper Freshwater-Molasse — 47.67°N, 8.93°EOler — 69.50°N, 158.50°EScarborough — 43.70°N, 79.24°WScarborough — 43.67°N, 79.32°WTitusville Till — 41.62°N, 79.64°WOlympia beds — 47.66°N, 122.43°WNiveau du gypse d'Aix — 43.52°N, 5.45°ETsubusagawa — 33.40°N, 131.35°EShanwang — 36.55°N, 118.71°ESalt — 47.69°N, 7.53°ENogami — 33.21°N, 131.15°EGreen River — 39.46°N, 108.41°WGreen River — 39.60°N, 108.40°WGreen River — 41.55°N, 109.51°WImuruk Volcanics — 65.88°N, 163.13°WOokui — 36.34°N, 138.34°EHirabaru — 33.40°N, 131.64°ESaijo — 34.38°N, 132.72°EMizozono — 31.98°N, 130.74°ECastle Eden Fissure-Fill — 54.74°N, 1.33°WTsumori — 32.79°N, 130.86°EQuadra Sand — 49.29°N, 122.88°W

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The named rock units Harpalinae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 15 further formations hold fewer occurrences each and are not listed. 223 occurrences have no formation on record, which is common for older literature.

Harpalinae and its relatives

Pleistocene

Where Harpalinae and its relatives were recorded in the Pleistocene.

Found in the same rock

6,282 taxa

The Paleobiology Database records 198 sites for Harpalinae, and this reads all of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.

This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.

Entered from

95 references

Each of the 548 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 83 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.

Media and terms

Media content1

Picture of Harpalinae

Credit
nobody named on the record
Licence
not stated by the source
Commercial use
cannot be read from the licence given — check the record

No 3D scan of Harpalinae was found in Sketchfab.

Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.

The description on this site

Licence
CC BY-SA 4.0

This is what the site found, which is not the same as what exists.

Keep going

What may have been alongside itCarabidae through timeFormations of the PleistoceneThe world it lived on

Occurrence data from the Paleobiology Database, CC BY 4.0. Bonelli 1810