Cytilus
Cytilus is a genus of pill beetles in the family Byrrhidae. There are about eight described species in Cytilus.

Ryan Hodnett · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- ByrrhiniPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 33 million years on record.
The record covers at least 33.9 Mya to 11.7 ka. No occurrence read is dated outside 37 Mya to 0 Mya.
15 occurrences of Cytilus on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
15 occurrences- Eocene7%
- Pleistocene87%
- Holocene7%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
4 formations
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The named rock units Cytilus has been recovered from, largest first. Drawn where the rock is now, not where it formed. 6 occurrences have no formation on record, which is common for older literature.
Cytilus and its relatives
PleistoceneWhere Cytilus and its relatives were recorded in the Pleistocene.
Found in the same rock
524 taxa- Stenus11 sites
- Bembidion10 sites
- Aleocharinae9 sites
- Bledius8 sites
- Helophorus8 sites
- Dyschirius7 sites
- Elateridae7 sites
- Notaris aethiops7 sites
- Quedius7 sites
- Aphodius6 sites
- Ochthebius6 sites
- Olophrum consimile6 sites
- Pselaphinae6 sites
- Aegialia5 sites
- Apion5 sites
- Catops5 sites
- Chrysomelidae5 sites
- Formicidae5 sites
- Omaliinae5 sites
- Patrobus stygicus5 sites
- Scydmaeninae5 sites
- Trichoptera5 sites
- Acidota crenata4 sites
- Arpedium cribratum4 sites
The Paleobiology Database records 13 sites for Cytilus, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
8 references- Fossil Beetle Evidence for a Short Warm Interval near 40,000 yr B.P. at Titusville, Pennsylvania4
- The paleoenvironment of the Olympia beds based on fossil beetles from Discovery Park, Seattle, Washington, U.S.A.3
- Palaeoenvironmental interpretation of Middle and Late Wisconsinan fossil coleopteran assemblages from western Olympic Peninsula, Washington, USA3
- Pill beetles (Coleoptera, Byrrhidae) in late Pleistocene deposits in the south of the west Siberia1
- Pre-Vashon fossil Coleoptera of Fraser age from the Fraser Lowland, British Columbia1
- The fossil Coleoptera of the Two Creeks Forest Bed, Wisconsin1
- A Late Pleistocene fauna and flora from Upton Warren, Worcestershire1
- Adephagous and clavicorn Coleoptera from the Tertiary deposits at Florissant, Colorado with descriptions of a few other forms and a systematic list of the non-rhynchophorus Tertiary Coleoptera of North America1
Each of the 15 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Cytilus
- Source
- Wikipedia/Commons
- Credit
- Ryan Hodnett
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan was found. Smithsonian Open Access, MorphoSource and Sketchfab were all searched.
The description on this site
- Source
- Wikipedia, “Cytilus”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itByrrhini through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Erichson 1847