Viperinae
Viperinae, or viperines, are a subfamily of vipers endemic to Europe, Asia and Africa. They are distinguished by their lack of the heat-sensing pit organs that characterize their sister group, the subfamily Crotalinae. Currently, 13 genera are recognized. Most are tropical and subtropical, although one species, Vipera berus, even occurs within the Arctic Circle. Like all vipers, they are venomous.

Uncredited · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- CrotalidaePBDB
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Amniota
Time the record covers
At least 22 million years on record.
The record covers at least 22.4 Mya to 11.7 ka. No occurrence read is dated outside 23.04 Mya to 0 Mya.
193 occurrences of Viperinae on record, most of them in the Miocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
193 occurrences- Miocene49%
- Pliocene20%
- Pleistocene31%
- Holoceneunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
24 formations
- Upper Freshwater Molasse Formation · Molasse Group6
- Vogel River Series Formation5
- Olduvai Formation3
- Cromer Forest Bed Formation2
- Akzhar Formation2
- Edelény Formation2
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The named rock units Viperinae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 152 occurrences have no formation on record, which is common for older literature.
Viperinae and its relatives
MioceneWhere Viperinae and its relatives were recorded in the Miocene.
Found in the same rock
1,780 taxa- Vipera79 sites
- Lacerta54 sites
- Rana40 sites
- Lacertidae39 sites
- Bufo bufo34 sites
- Colubridae32 sites
- Natrix31 sites
- Anguis fragilis30 sites
- Ophisaurus28 sites
- Pseudopus27 sites
- Colubrinae26 sites
- Natrix natrix23 sites
- Vipera (Pelias) berus23 sites
- Natricinae22 sites
- Serpentes22 sites
- Coluber20 sites
- Pelobates20 sites
- Elaphe19 sites
- Bufo18 sites
- Bufo viridis18 sites
- Coronella austriaca18 sites
- Gekkonidae17 sites
- Hyla arborea17 sites
- Pelophylax17 sites
The Paleobiology Database records 155 sites for Viperinae, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
82 references- Pleistocene Amphibians and Reptiles in Britain and Europe21
- Late Cainozoic localities of snakes and lizards of Ukraine15
- Oldest Fossil Vipers (Serpentes: Viperidae) from the Old World11
- Fossil snakes from Poland11
- Amphibians and squamate reptiles from the late Miocene of Fălciu (Eastern Romania)6
- Amphibians and Squamates from the Miocene of Felsőtárkány Basin, N-Hungary5
- Long-term climate record inferred from early-middle Pleistocene amphibian and squamate reptile assemblages at the Gran Dolina Cave, Atapuerca, Spain5
- Fossil snake assemblages from the French Middle Miocene localities at La Grive (France)5
- Fossil snake localities of the Caucasus5
- Vipera (Daboia) ukrainica – a new viper (Serpentes; Viperidae) from the Middle Sarmatian (Upper Miocene) of the Ukraine5
- Laetoli: a Pliocene Site in Northern Tanzania5
- Snakes from Griesbeckerzell (Langhian, Early Badenian), North Alpine Foreland Basin (Germany), with comments on the evolution of snake faunas in Central Europe during the Miocene Climatic Optimum3
Each of the 193 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 70 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Viperinae
- Source
- Wikipedia/Commons
- Credit
- nobody named on the record
- Licence
- not stated by the source
- Commercial use
- cannot be read from the licence given — check the record
No 3D scan of Viperinae was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Viperinae”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itCrotalidae through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Oppel 1811)