Testudo graeca
The Greek tortoise, also known commonly as the Moorish tortoise and the spur-thighed tortoise, is a species of tortoise in the family Testudinidae. The species is a medium-sized herbivorous testudinid, widely distributed in the Mediterranean region.

Charles J. Sharp · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- common tortoisePBDB
- Sits inside
- TestudoPBDB
- Diet
- herbivore, herbivorePBDB, for Testudinidae
- Life habit
- ground dwellingPBDB, for Testudinidae
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Testudinidae
Time the record covers
At least 11 million years on record.
The record covers at least 11.63 Mya to 11.7 ka. No occurrence read is dated outside 15.98 Mya to 0 Mya.
32 occurrences of Testudo graeca on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
32 occurrences- Miocene39%
- Pliocene11%
- Pleistocene41%
- Holocene9%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
1 formation
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The named rock units Testudo graeca has been recovered from, largest first. Drawn where the rock is now, not where it formed. 31 occurrences have no formation on record, which is common for older literature.
Testudo graeca and its relatives
PleistoceneWhere Testudo graeca and its relatives were recorded in the Pleistocene.
Found in the same rock
284 taxa- Equus4 sites
- Gazella4 sites
- Gerbillus4 sites
- Meriones4 sites
- Allocricetus3 sites
- Apodemus3 sites
- Bos primigenius3 sites
- Bufo viridis3 sites
- Chiroptera3 sites
- Crocidurinae3 sites
- Emys orbicularis3 sites
- Lacerta viridis3 sites
- Lepus capensis3 sites
- Mus musculus3 sites
- Oryctolagus3 sites
- Panthera leo3 sites
- Rhinolophus3 sites
- Rhinolophus ferrumequinum3 sites
- Soricinae3 sites
- Spalax3 sites
- Allohippus stenonis2 sites
- Arvicola2 sites
- Bufo2 sites
- Bufo bufo2 sites
The Paleobiology Database records 31 sites for Testudo graeca, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
15 references- Fossile Schildkroten aus der Turkei12
- The Fossil Chelonians of Greece. Systematics - Evolution - Stratigraphy - Palaeoecology3
- The oldest evidence of Testudo graeca (Testudinidae) in the Iberian Peninsula2
- The microvertebrates of Qesem Cave: A comparison of the two concentrations2
- The chelonians from the Latest Miocene–Earliest Pliocene localities of Allatini and Pylea (East Thessaloniki, Macedonia, Greece)2
- Pleistocene Amphibians and Reptiles in Britain and Europe2
- Freshwater turtle or tortoise? The exploitation of testudines at the Mousterian site of Nahal Mahanayeem Outlet, Hula Valley, Israel1
- Paleoclimatic and paleoenvironmental context of the Early Pleistocene hominins from Dmanisi (Georgia, Lesser Caucasus) inferred from the herpetofaunal assemblage1
- Testudo cf. graeca from the new Late Miocene locality of Platania (Drama basin, N. Greece) and a reappraisal of previously published specimens1
- Outside Africa: Middle Pleistocene Lycaon from Hayonim Cave, Israel1
- Kupferzeitliche Schildkrötenreste aus der Grabung von Durankulak bei Tolbuchin in NO-Bulgarien (Reptilia: Testudines: Testudo graeca, Emys orbicularis)1
- The Mousterian site of Jebel Irhoud, Morocco: new data on the fauna and biochronology, and description of a new human fossil1
Each of the 32 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 3 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Testudo graeca
- Source
- Wikipedia/Commons
- Credit
- Charles J. Sharp
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan of Testudo graeca was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itTestudo through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Linnaeus 1758