Carabus
Carabus is a genus of beetles in the family Carabidae, and was first described in 1758 by Carl Linnaeus.

Dodoni · CC BY 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- CarabiniPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 40 million years on record.
The record covers at least 41.03 Mya to 129 ka. No occurrence read is dated outside 48.07 Mya to 11.7 ka.
51 occurrences of Carabus on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
51 occurrences- Eocene6%
- Miocene14%
- Pliocene2%
- Pleistocene78%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
17 formations
- Titusville Till Formation5
- Bushi Formation3
- Ookui Formation3
- Hirabaru Formation3
- Nogami Formation3
- Shanwang Formation2
Loading coastlines…
The named rock units Carabus has been recovered from, largest first. Drawn where the rock is now, not where it formed. 20 occurrences have no formation on record, which is common for older literature.
Carabus and its relatives
PleistoceneWhere Carabus and its relatives were recorded in the Pleistocene.
Found in the same rock
1,188 taxa- Bembidion14 sites
- Aphodius12 sites
- Carabidae12 sites
- Agabus10 sites
- Coleoptera9 sites
- Stenus9 sites
- Curculionidae8 sites
- Elateridae8 sites
- Pterostichus (Cryobius)8 sites
- Staphylinidae8 sites
- Hydrobius fuscipes7 sites
- Ilybius7 sites
- Olophrum rotundicolle7 sites
- Phyllobius7 sites
- Tournotaris bimaculatus7 sites
- Cholevinus sibiricus6 sites
- Chrysomelidae6 sites
- Curtonotus alpinus6 sites
- Donacia6 sites
- Dyschirius6 sites
- Gyrinus6 sites
- Helophorus6 sites
- Hypera ornata6 sites
- Morychus viridis6 sites
The Paleobiology Database records 39 sites for Carabus, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
27 references- Late Cenozoic insects of northern Eurasia6
- Fossil Beetle Evidence for a Short Warm Interval near 40,000 yr B.P. at Titusville, Pennsylvania5
- Pleistoceńskie owady z Borysławia4
- Middle Pleistocene fossil insects from the Nogami Formation of Kokonoe, Oita Prefecture, Kyushu, Japan3
- Middle Pleistocene fossil insects from the Hirabaru Formation of Kitsuki City, Oita Prefecture, Kyushu, Japan3
- Early Pleistocene ground beetles (Coleoptera: Carabidae) from the Ookui Formation in Nagano Prefecture, central Japan, and their biogeographical and paleoenvironmental significance3
- Insect fossil assemblage and paleoenvironments from the Early Pleistocene Bushi Formation in Saitama Prefecture, Japan3
- The paleoenvironment of the Olympia beds based on fossil beetles from Discovery Park, Seattle, Washington, U.S.A.2
- Stratigraphy, fossils, and age of sediments at the upper pit of the Lost Chicken gold mine: new information on the late Pliocene environment of east central Alaska2
- Trois fossiles remarquablement conservés du Miocène de France, appartenant aux genres Carabus L. et Ledouxnebria nov. (Coleoptera, Carabidae et Nebriidae)2
- A Late Pleistocene fauna and flora from Upton Warren, Worcestershire2
- Pleistocene fossil insects from Tokachi, Hokkaido, north Japan1
Each of the 51 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 15 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content3
Picture of Carabus
- Source
- Wikipedia/Commons
- Credit
- Dodoni
- Licence
- CC BY 3.0
- Commercial use
- the licence allows it
Photogrammetry
- Source
- MorphoSource
- Licence
- not stated by the source
- Commercial use
- cannot be read from the licence given — check the record
- Specimen
- Carabus Carabus catenulatus · ETHZ:ENT:0577357
3D model
- Source
- Sketchfab
- Licence
- CC Attribution
- Commercial use
- cannot be read from the licence given — check the record
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Carabus”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itCarabini through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Linnaeus 1758