Scaphopoda
Scaphopoda is a class of shelled marine molluscs, whose members are known as scaphopods and commonly called tusk shells or tooth shells. They have a worldwide distribution and are the only class of exclusively infaunal marine molluscs. Shells of species within this class range in length 0.5–18 cm (0.20–7.09 in), with Fissidentalium metivieri being the longest. Members of the order Dentaliida tend to be larger than those of the order Gadilida.

[Image: Glover, Dahlgren & Wiklund, 2017] Wiklund H, Taylor JD, Dahlgren TG, Todt C, Ikebe C, Rabone M, Glover AG (20… · CC BY 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- tusk shellPBDB
- Sits inside
- MolluscaPBDB
- Diet
- deposit feederPBDB
- Life habit
- shallow infaunalPBDB
- Motility
- slow-movingPBDB
- Vision
- blindPBDB
Time the record covers
At least 422 million years on record.
The record covers at least 422.7 Mya to 11.7 ka. No occurrence read is dated outside 426.7 Mya to 0 Mya.
3,856 occurrences of Scaphopoda on record, most of them in the Miocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
3,856 occurrences- Silurianunder 1%
- Devonianunder 1%
- Carboniferous3%
- Permian6%
- Early Triassicunder 1%
- Middle Triassic1%
- Late Triassic2%
- Early Jurassic3%
- Middle Jurassic3%
- Late Jurassicunder 1%
- Early Cretaceous1%
- Late Cretaceous11%
- Paleocene6%
- Eocene19%
- Oligocene8%
- Miocene20%
- Pliocene9%
- Pleistocene6%
- Holoceneunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
581 formations
- Gatun Formation117
- Cook Mountain Formation · Claiborne Group110
- Moodys Branch Formation · Jackson Group74
- Corsicana Marl Formation69
- Providence Formation61
- Cassian Formation52
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The named rock units Scaphopoda has been recovered from, largest first. Drawn where the rock is now, not where it formed. 557 further formations hold fewer occurrences each and are not listed. 687 occurrences have no formation on record, which is common for older literature.
Scaphopoda and its relatives
MioceneWhere Scaphopoda and its relatives were recorded in the Miocene.
Found in the same rock
2,164 taxa- Dentalium107 sites
- Cadulus88 sites
- Gastropoda70 sites
- Turritella67 sites
- Nuculana57 sites
- Corbula45 sites
- Bivalvia40 sites
- Calyptraea (Trochita) aperta37 sites
- Bryozoa36 sites
- Naticidae36 sites
- Hipponix35 sites
- Vertebrata34 sites
- Syncyclonema33 sites
- Nucula32 sites
- Retusa galba32 sites
- Anthozoa31 sites
- Athleta petrosa31 sites
- Eburneopecten scintillatus31 sites
- Striarca30 sites
- Anomia argentaria28 sites
- Cyprimeria28 sites
- Acteon27 sites
- Bullata semen27 sites
- Endopachys maclurii27 sites
The Paleobiology Database records at least 1,410 sites for Scaphopoda, and this reads 240 of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
663 references- Resilience of marine invertebrate communities during the early Cenozoic hyperthermals158
- Die Fauna des marinen Miozäns von Kevelaer (Niederrhein). Band 2 Bivalvia, Scaphopoda, Cephalopoda, Bryozoa, Annelida, Brachiopoda148
- A paleobiotic survey of Caribbean faunas from the Neogene of the Isthmus of Panama113
- Catalogue of the Paleocene and Eocene mollusca of the southern and eastern United States. Part 1. Pelecypoda, Amphineura, Pteropoda, Scaphopoda and Cephalopoda100
- Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (additions to gastropods, scaphopods, pelecypods: Nuculidae to Malleidae)97
- Palökologie von Benthos-Gemeinschaften im oberen Pliensbach (Amaltheenton) von Kalchreuth (Mittelfranken)76
- Faunal dynamics of bivalves and scaphopods in the Bathonian (Middle Jurassic) ore-bearing clays at Gnaszyn, Kraków-Silesia Homocline, Poland73
- Upper Cretaceous (Maestrichtian) larger invertebrate Fossils from the Haustator bilira Assemblage Zone in the west gulf Coastal Plain72
- Upper Cretaceous (Maestrichtian) larger invertebrates from the Haustator bilira Assemblage Zone in the Atlantic Coastal Plain with further data for the East Gulf62
- Unpublished notes on Caribbean collection of the Naturhistorisches Museum Basel59
- Upper Cretaceous (Maestrichtian) Mollusca from the Haustator bilira Assemblage Zone in the East Gulf Coastal Plain46
- The Molluscan Fauna of the Alum Bluff Group of Florida44
Each of the 3,856 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 651 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Scaphopoda
- Source
- Wikipedia/Commons
- Credit
- [Image: Glover, Dahlgren & Wiklund, 2017] Wiklund H, Taylor JD, Dahlgren TG, Todt C, Ikebe C, Rabone M, Glover AG (20…
- Licence
- CC BY 4.0
- Commercial use
- the licence allows it
No 3D scan of Scaphopoda was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Tusk shell”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itMollusca through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Bronn 1862