Pugnoides
Pugnoides is an extinct genus of brachiopod belonging to the order Rhynchonellida and family Petasmariidae. Specimens have been found in Devonian to Permian beds in North America, Asia, Europe, western Australia, New Zealand,and New Zealand. The genus was particularly widespread in the Visean.

via iDigBio · CC0 · via iDigBio. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- PetasmariidaePBDB
- Diet
- suspension feederPBDB, for Rhynchonellida
- Life habit
- epifaunalPBDB, for Rhynchonellida
- Motility
- stationaryPBDB, for Rhynchonellida
- Vision
- blindPBDB, for Rhynchonellata
- Environment
- marinePBDB, for Lophophorata
Time the record covers
At least 118 million years on record.
The record covers at least 382.31 Mya to 264.28 Mya. No occurrence read is dated outside 387.95 Mya to 259.857 Mya.
63 occurrences of Pugnoides on record, most of them in the Carboniferous. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
63 occurrences- Devonian40%
- Carboniferous43%
- Permian17%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
28 formations
- Triangle Formation · Sonyea Group14
- Amsden Formation10
- Kaibab Formation3
- Otter Formation2
- Lime Creek Formation2
- Panther Seep Formation2
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The named rock units Pugnoides has been recovered from, largest first. Drawn where the rock is now, not where it formed. 4 further formations hold fewer occurrences each and are not listed. 6 occurrences have no formation on record, which is common for older literature.
Pugnoides and its relatives
CarboniferousWhere Pugnoides and its relatives were recorded in the Carboniferous.
Found in the same rock
834 taxa- Crinoidea23 sites
- Leiorhynchus18 sites
- Productella17 sites
- Schizophoria16 sites
- Composita15 sites
- Eoschizodus15 sites
- Bryozoa14 sites
- Strophomena (Strophodonta)13 sites
- Mucrospirifer12 sites
- Palaeoneilo constricta11 sites
- Atrypa10 sites
- Spirifer10 sites
- Nautiloidea9 sites
- Cyrtina8 sites
- Eumetria8 sites
- Actinopteria7 sites
- Camarotoechia7 sites
- Chonetes7 sites
- Gastropoda7 sites
- Warrenella7 sites
- Bellerophontoidea6 sites
- Composita sulcata6 sites
- Lyriopecten6 sites
- Plantae6 sites
The Paleobiology Database records 61 sites for Pugnoides, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
29 references- Quantitative community analysis: Recurrent group and cluster techniques applied to the fauna of the Upper Devonian Sonyea Group, New York15
- Stratigraphy and geologic history of the Amsden Formation (Mississippian and Pennsylvanian) of Wyoming10
- The Guadalupian fauna4
- The environment and history of the Toroweap and Kaibab Formations of northern Arizona and southern Utah3
- Homeomorphy between Devonian brachiopod communities in France and Iowa2
- Marine paleoecology in the Upper Devonian of New York2
- Geology of the Bear Park Area, Dona Ana County, New Mexico2
- Biostratigraphy and Carbonate Facies of the Mississippian Arroyo Penasco Formation, North - Central New Mexico2
- Carboniferous formations and faunas of central Montana2
- The Paradise Formation and its fauna2
- Work with Thayer collections, Yale Peabody Accession 7112, published, in part, "New perspectives on transitions between ecological-evolutionary subunits in the "type interval" for coordinated stasis", James J Zambito IV, Carlton E Brett, Gordon C Baird, Sarah E Kolbe, Arnold I Miller; Paleobiology1
- Silicified Carboniferous (Chesterian) Brachiopoda of the Arco Hills Formation, Idaho1
Each of the 63 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 17 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1

Picture of Pugnoides
- Source
- iDigBio
- Credit
- via iDigBio
- Licence
- CC0
- Commercial use
- the licence allows it
No 3D scan was found. Smithsonian Open Access, MorphoSource and Sketchfab were all searched.
The description on this site
- Source
- Wikipedia, “Pugnoides”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itPetasmariidae through timeFormations of the CarboniferousThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Weller 1910)