Ilex aquifolium
Ilex aquifolium, the holly, common holly, English holly, European holly, or occasionally Christmas holly, is a species of flowering plant in the family Aquifoliaceae, native to western and southern Europe, northwest Africa, and southwest Asia. It is regarded as the type species of the genus Ilex, which by association is also called holly. It is an evergreen tree or shrub found, for example, in shady areas of forests of oak and in beech hedges. In the British Isles it is one of very few native hardwood evergreen trees. It has a great capacity to adapt to different conditions and is a pioneer species that repopulates the margins of forests or clearcuts.

Jürgen Howaldt · CC BY-SA 2.0 de · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- IlexPBDB
- Diet
- "photoautotroph"PBDB, for Angiospermae
Time the record covers
At least 8 million years on record.
The record covers at least 11.63 Mya to 3.6 Mya. No occurrence read is dated outside 15.98 Mya to 11.7 ka.
16 occurrences of Ilex aquifolium on record, most of them in the Pliocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
16 occurrences- Miocene16%
- Pliocene81%
- Pleistocene3%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
1 formation
Loading coastlines…
The named rock units Ilex aquifolium has been recovered from, largest first. Drawn where the rock is now, not where it formed. 15 occurrences have no formation on record, which is common for older literature.
Ilex aquifolium and its relatives
PlioceneWhere Ilex aquifolium and its relatives were recorded in the Pliocene.
Found in the same rock
119 taxa- Acer10 sites
- Quercus10 sites
- <not identified>9 sites
- Alnus9 sites
- Cathaya9 sites
- Cistaceae9 sites
- Engelhardia9 sites
- Ericaceae9 sites
- Myrica9 sites
- Pinus9 sites
- Asteraceae8 sites
- Juglans8 sites
- Oleaceae8 sites
- Osmunda8 sites
- Poaceae8 sites
- Taxodiaceae8 sites
- <Amaranthaceae/Chenopodiaceae>7 sites
- Apiaceae7 sites
- Betula7 sites
- Carya7 sites
- Castanea7 sites
- Cupressaceae7 sites
- Gleicheniaceae7 sites
- Nyssa7 sites
The Paleobiology Database records 10 sites for Ilex aquifolium, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
7 references- Niche evolution versus niche conservatism and habitat loss determine persistence and extirpation in late Neogene European Fagaceae9
- Die pliozaenen Floren von Thueringen, Deutsche Demokratische Republik2
- The section of Torre Picchio (Terni, Umbria, Central Italy): A Villafranchian site rich in vertebrates, molluscs, ostracods and plants.1
- Die mittelmiozaenen und obermiozaenen Floren aus der Meuroer und Raunoer Folge in der Lausitz. III. Fundstellen und Palaeobiologie1
- Miocene floras in the lower Rhenish basin and their ecological interpretation1
- Allochthonous seed and fruit floras from the Pliocene of the Lower Rhine Basin1
- Miocenska Flora ze Starych Gliwic na Slasku (Miocene Flora of Stare Gliwice in Upper Silesia)1
Each of the 16 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Ilex aquifolium
- Source
- Wikipedia/Commons
- Credit
- Jürgen Howaldt
- Licence
- CC BY-SA 2.0 de
- Commercial use
- the licence allows it
No 3D scan of Ilex aquifolium was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itIlex through timeFormations of the PlioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Linnaeus 1753