Cyphaspis
Cyphaspis is a genus of small, proetid trilobite that lived from the Late Ordovician to the Late Devonian. Fossils have been found in marine strata in what is now Europe, Africa and North America. Various species had a compact body, and a large, bulbous glabellum. Many species had long spines arranged similarly to closely related genera, such as Otarian, Otarionella, Chamaeleoaspis, and Namuropyge.

Jonathan Chen · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- OtarioniniPBDB
- Diet
- deposit feederPBDB, for Proetida
- Life habit
- low-level epifaunalPBDB, for Trilobita
- Motility
- fast-movingPBDB, for Trilobita
- Environment
- marinePBDB, for Trilobita
Time the record covers
At least 96 million years on record.
The record covers at least 471.3 Mya to 375.2 Mya. No occurrence read is dated outside 475.3 Mya to 372.15 Mya.
128 occurrences of Cyphaspis on record, most of them in the Devonian. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
128 occurrences- Ordovician9%
- Silurian10%
- Devonian81%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
57 formations
- Jemelle Formation7
- Timrhanrhart Formation6
- Zlichov Limestone Formation5
- Ihandar Formation4
- Ahrdorf Formation4
- Chefar el Ahmar Formation4
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The named rock units Cyphaspis has been recovered from, largest first. Drawn where the rock is now, not where it formed. 33 further formations hold fewer occurrences each and are not listed. 26 occurrences have no formation on record, which is common for older literature.
Cyphaspis and its relatives
DevonianWhere Cyphaspis and its relatives were recorded in the Devonian.
Found in the same rock
1,559 taxa- Leonaspis18 sites
- Crotalocephalina gibbus13 sites
- Koneprusia13 sites
- Ostracoda12 sites
- Scutellum12 sites
- Tropidocoryphe12 sites
- Ceratocephala11 sites
- Crinoidea11 sites
- Nowakia11 sites
- Dicranurus monstrosus10 sites
- Scabriscutellum10 sites
- Gastropoda9 sites
- Reedops bronni9 sites
- Zlichovaspis (Zlichovaspis) rugosa9 sites
- Atrypa8 sites
- Crotalocephalus8 sites
- Favosites8 sites
- Platyscutellum8 sites
- Polygnathus8 sites
- Reedops cephalotes8 sites
- Reedops sternbergi8 sites
- Cornuproetus7 sites
- Nucleospira7 sites
- Proetus7 sites
The Paleobiology Database records 97 sites for Cyphaspis, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
52 references- Trilobite assemblages in the Devonian of the Barrandian area and their relations to palaeoenvironments14
- The Bohemian Lower Devonian Stages13
- Morphological trends and new species of Cyphaspis (Trilobita, Otarioninae) in the Devonian of Morocco, Turkey, Germany and Belgium8
- Eifel-Trilobiten III. Corynexochida, Proetida (2), Harpetida, Phacopida (2), Lichida6
- Revision der Ostbaltischen Silurichen Trilobiten. Abtheilung IV: Calymmeniden, Proetiden, Bronteiden, Harpediden, Trunicleiden, Remopleuriden und Agnostiden.6
- Trilobites from the upper Lower to Middle Devonian Timrhanrhart Formation, Jbel Gara el Zguilma, southern Morocco5
- The phacopid trilobites Austerops McKellar & Chatterton, 2009, Hottonops gen. nov. and Loreleiops gen. nov. from the Devonian of the Ardenno-Rhenish Mountains4
- A trilobite faunule from the Lower Devonian of the Saoura Valley, Algeria: biodiversity, morphological variability and palaeobiogeographical affinities4
- Proetid Trilobites from the Lower Devonian (Pragian) Ihandar Formation, Anti-Atlas, Morocco4
- The rise and fall of Late Devonian (Frasnian) trilobites from Belgium: taxonomy, biostratigraphy and events3
- “Devil horned” Cyphaspis (Trilobita, Otarioninae): examples from the Middle Devonian of the Ardennes (Belgium), Eifel (Germany) and Ma’der (Morocco)3
- Taxonomy and biostratigraphy of some proetid trilobites in the Middle Devonian of the Ardennes and Eifel (Rhenohercynian Zone)3
Each of the 128 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 40 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Cyphaspis
- Source
- Wikipedia/Commons
- Credit
- Jonathan Chen
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan of Cyphaspis was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Cyphaspis”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itOtarionini through timeFormations of the DevonianThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Burmeister 1843)