Dryopidae
Dryopidae is a family of beetles, commonly named long-toed water beetles, in the superfamily Byrrhoidea. It was described by Gustaf Johan Billberg in 1820.

S.E. Thorpe · Public Domain · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- long toed water beetlePBDB
- Sits inside
- DryopoideaPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 33 million years on record.
The record covers at least 33.9 Mya to 129 ka. No occurrence read is dated outside 37 Mya to 11.7 ka.
31 occurrences of Dryopidae on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
31 occurrences- Eocene13%
- Oligocene3%
- Miocene3%
- Pleistocene81%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
3 formations
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The named rock units Dryopidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 14 occurrences have no formation on record, which is common for older literature.
Dryopidae and its relatives
PleistoceneWhere Dryopidae and its relatives were recorded in the Pleistocene.
Found in the same rock
739 taxa- Aphodius16 sites
- Stenus16 sites
- Aleocharinae14 sites
- Dryops14 sites
- Dyschiriodes globosus14 sites
- Anotylus rugosus13 sites
- Hydraena13 sites
- Rhynchaenus quercus13 sites
- Oulimnius troglodytes12 sites
- Apion11 sites
- Bryaxis11 sites
- Esolus parallelepipedus11 sites
- Limnius volckmari11 sites
- Oulimnius tuberculatus11 sites
- Helodidae10 sites
- Otiorhynchus (Pendragon) ovatus10 sites
- Bagous9 sites
- Bembidion (Diplocampa) assimile9 sites
- Carpelimus9 sites
- Gyrinus9 sites
- Ochthebius minimus9 sites
- Pelochares versicolor9 sites
- Adelocera murina8 sites
- Elmis aenea8 sites
The Paleobiology Database records 24 sites for Dryopidae, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
12 references- Marine Isotope Stage 9 environments of fluvial deposits at Hackney, north London, UK12
- Middle Pleistocene deposits at Frog Hall Pit, Stretton-on-Dunsmore, Warwickshire, English Midlands, and their implications for the age of the type Wolstonian5
- Late Devensian river deposits beneath the floodplain terrace of the River Thames at Abingdon, Berkshire, England4
- New Miocene Coleoptera from Florissant2
- Paleogeography of the lower Volga at the end of Atelian time based on entomological and botanical data1
- Fossile Käfer aus miozänen Sedimenten des Randecker Maars in Südwest-Deutschland (Insecta: Coleoptera)1
- A Late Pleistocene fauna and flora from Upton Warren, Worcestershire1
- Miozäne und oligozäne Insekten aus Österreich und den unmittelbar angrenzenden Gebieten1
- Note Complémentaire sur les insectes fossiles des Cinérites Pliocènes du Lac Chambon (Puy-de-Dôme)1
- A report on some recent collections of fossil Coleoptera from the Miocene shales of Florissant1
- Fossil Coleoptera from Florissant, with descriptions of several new species1
- Pleistoceńskie owady z Borysławia1
Each of the 31 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Dryopidae
- Source
- Wikipedia/Commons
- Credit
- S.E. Thorpe
- Licence
- Public Domain
- Commercial use
- the licence allows it
No 3D scan of Dryopidae was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Dryopidae”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itDryopoidea through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Billberg 1820