Aquifoliaceae
Ilex or holly is a genus of over 570 species of flowering plants in the family Aquifoliaceae, and the only living genus in that family. Ilex has the most species of any woody dioecious angiosperm genus. The species are evergreen or deciduous trees, shrubs, and climbers from tropics to temperate zones worldwide. The type species is Ilex aquifolium, the common European holly used in Christmas decorations and cards.

Jürgen Howaldt · CC BY-SA 2.0 de · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- AquifolialesPBDB
- Diet
- "photoautotroph"PBDB, for Angiospermae
Time the record covers
At least 93 million years on record.
The record covers at least 93.9 Mya to 11.7 ka. No occurrence read is dated outside 143.1 Mya to 0 Mya.
300 occurrences of Aquifoliaceae on record, most of them in the Miocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
300 occurrences- Early Cretaceousunder 1%
- Late Cretaceous4%
- Paleocene8%
- Eocene24%
- Oligocene3%
- Miocene33%
- Pliocene14%
- Pleistocene10%
- Holocene2%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
56 formations
- Río Turbio Formation11
- Lefipán Formation8
- Yaw Formation8
- Gatun Formation6
- Culebra Formation5
- Itahana Formation · Tomioka Group4
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The named rock units Aquifoliaceae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 32 further formations hold fewer occurrences each and are not listed. 187 occurrences have no formation on record, which is common for older literature.
Aquifoliaceae and its relatives
MioceneWhere Aquifoliaceae and its relatives were recorded in the Miocene.
Found in the same rock
2,043 taxa- Ilex78 sites
- Ilexpollenites58 sites
- Poaceae49 sites
- Asteraceae43 sites
- Carya42 sites
- Alnus39 sites
- Podocarpidites38 sites
- Quercus35 sites
- Triatriopollenites35 sites
- Laevigatosporites34 sites
- Podocarpus34 sites
- Botryococcus33 sites
- Cyathea33 sites
- Momipites33 sites
- Betula32 sites
- Ericaceae32 sites
- Myrtaceae32 sites
- Pinus31 sites
- Symplocos31 sites
- Cyperaceae30 sites
- Nothofagidites30 sites
- Proteacidites30 sites
- Liquidambar29 sites
- Plicatopollis29 sites
The Paleobiology Database records 195 sites for Aquifoliaceae, and this reads all of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
77 references- Die mittelmiozaenen und obermiozaenen Floren aus der Meuroer und Raunoer Folge in der Lausitz. III. Fundstellen und Palaeobiologie29
- Vegetation dynamics in a disturbed lacustrine record: The Eocene maar lake of Groß-Zimmern (Hesse, Germany)18
- Allochthonous seed and fruit floras from the Pliocene of the Lower Rhine Basin16
- Early Pleistocene vegetation change in upland south-eastern Australia14
- Miocene floras in the lower Rhenish basin and their ecological interpretation13
- Early Tertiary palaeogeography, landform evolution, and palaeoclimates of the Southern Monaro, N.S.W., Australia12
- Material fúngico y granos de polen de angiospermas de la Formación Río Turbio (Eoceno), provincia de Santa Cruz, República Argentina12
- A Catalogue of Fossil Plants from India Part 6: Cenozoic (Quaternary) Palynology and Palaeobotany11
- Niche evolution versus niche conservatism and habitat loss determine persistence and extirpation in late Neogene European Fagaceae9
- Late Miocene and Pliocene Floras in Central Honshu, Japan9
- Stratigraphic, Paleoclimatic, and Paleobiogeographic Significance of Tertiary Sporomorphs from Massachusetts9
- Paleogene Sporomorph Biostratigraphy, Northeastern Virginia9
Each of the 300 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 65 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Aquifoliaceae
- Source
- Wikipedia/Commons
- Credit
- Jürgen Howaldt
- Licence
- CC BY-SA 2.0 de
- Commercial use
- the licence allows it
No 3D scan of Aquifoliaceae was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Holly”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itAquifoliales through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Richard 1820