Ampulicidae
Ampulicidae, or cockroach wasps, are a small, primarily tropical family of sphecoid wasps, all of which use various cockroaches as prey for their larvae. They are the most primitive family of sphecoid hunting wasps. They tend to have elongated jaws, pronounced neck-like constrictions behind the head, strongly petiolate abdomens, and deep grooves on the thorax. Many are quite ant-like in appearance, though some are brilliant metallic blue, green, and hot pink.

Muhammad Mahdi Karim · GFDL 1.2 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- cockroach waspPBDB
- Sits inside
- ApoideaPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 92 million years on record.
The record covers at least 113.2 Mya to 20.45 Mya. No occurrence read is dated outside 119.57 Mya to 15.98 Mya.
10 occurrences of Ampulicidae on record, most of them in the Early Cretaceous. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
10 occurrences- Early Cretaceous40%
- Late Cretaceous30%
- Eocene20%
- Miocene10%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
2 formations
- Crato Formation · Santana Group4
- Shanwang Formation1
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The named rock units Ampulicidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 5 occurrences have no formation on record, which is common for older literature.
Ampulicidae and its relatives
Early CretaceousWhere Ampulicidae and its relatives were recorded in the Early Cretaceous.
Found in the same rock
664 taxa- Mariconitermes talicei3 sites
- Meiatermes araripena3 sites
- Araripeneura damianii2 sites
- Araripeneura regia2 sites
- Cariridris bipetiolata2 sites
- Caririlabia santanensis2 sites
- Cearagrylloides perforatorius2 sites
- Chresmoda neotropica2 sites
- Colocrus indivicum2 sites
- Cordulagomphus fenestratus2 sites
- Cratoelcana zessini2 sites
- Cratomastotermes wolfschwenningeri2 sites
- Cretereisma2 sites
- Cretolabia cearae2 sites
- Cupedidae2 sites
- Elateridae2 sites
- Elisama brevis2 sites
- Eoprotoneura hyperstigma2 sites
- Ephemeroidea2 sites
- Ephemeroptera2 sites
- Gomphaeschnaoides obliquus2 sites
- Kotejalabis goethitica2 sites
- Leptophlebiidae2 sites
- Leptotarsus grimaldii2 sites
The Paleobiology Database records 8 sites for Ampulicidae, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
8 references- Martill et al., 20072
- Digger wasps (Hymenoptera, Sphecidae) in Burmese amber2
- The First Fossil Representative of the Wasp Genus Dolichurus, with a Review of Fossil Ampulicidae (Hymenoptera: Apoidea)1
- The earliest known fossil ant (first southern hemisphere Mesozoic record) (Hymenoptera: Formicidae: Myrmeciinae)1
- Insects from the Santana Formation, Lower Cretaceous, of Brazil. Chapter 7: Order Hymenoptera1
- Zhang, 19891
- A new genus of digger wasps from the Baltic amber1
- Zur Systematik und Palökologie harzkonservierter Arthropoda einer Taphozönose aus dem Cenomanium von NW-Frankreich1
Each of the 10 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Ampulicidae
- Source
- Wikipedia/Commons
- Credit
- Muhammad Mahdi Karim
- Licence
- GFDL 1.2
- Commercial use
- the licence allows it
No 3D scan of Ampulicidae was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Ampulicidae”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itApoidea through timeFormations of the Early CretaceousThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Shuckard 1840