Polyxenida
Polyxenida is an order of millipedes readily distinguished by a unique body plan consisting of a soft, non-calcified body ornamented with tufts of bristles. These features have inspired the common names bristly millipedes or pincushion millipedes. This order includes about 148 species in four families worldwide, which represent the only living members of the subclass Penicillata.

Sarefo · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- PenicillataPBDB
- Motility
- actively mobilePBDB, for Arthropoda
Time the record covers
At least 100 million years on record.
The record covers at least 121.4 Mya to 20.45 Mya. No occurrence read is dated outside 125.77 Mya to 13.82 Mya.
23 occurrences of Polyxenida on record, most of them in the Late Cretaceous. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
23 occurrences- Early Cretaceous13%
- Late Cretaceous48%
- Eocene26%
- Miocene13%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
1 formation
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The named rock units Polyxenida has been recovered from, largest first. Drawn where the rock is now, not where it formed. 22 occurrences have no formation on record, which is common for older literature.
Polyxenida and its relatives
Late CretaceousWhere Polyxenida and its relatives were recorded in the Late Cretaceous.
Found in the same rock
2,450 taxa- Araneae4 sites
- Phryssonotus burmiticus4 sites
- Geophilomorpha3 sites
- Mygalomorphae3 sites
- Polydesmida3 sites
- Polyxenidae3 sites
- Polyzoniida3 sites
- Scolopendromorpha3 sites
- Siphonophoridae3 sites
- Ablator triguttatus2 sites
- Adorator hispidus2 sites
- Aleyrodidae2 sites
- Allacrotelsa dubia2 sites
- Andrognathus burmiticus2 sites
- Anyphaena fuscata2 sites
- Archaea paradoxa2 sites
- Burmadictyna excavata2 sites
- Burmathele2 sites
- Burmesarchaea bilongapophyses2 sites
- Burmesarchaea pseudogibber2 sites
- Burmorchestina pulchra2 sites
- Burmorsolus2 sites
- Caddo dentipalpus2 sites
- Cataglyphoides constricta2 sites
The Paleobiology Database records 14 sites for Polyxenida, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
10 references- Checklist of the Myriapoda in Cretaceous Burmese amber and a correction of the Myriapoda identified by Zhang (2017)6
- Footnotes and taxonomic names, in Die im Bernstein befindlichen Crustaceen, Myriapoden, Arachniden und Apteren der Vorwelt4
- The oldest records of Polyxenida (Myriapoda, Diplopoda): new discoveries from the Cretaceous ambers of Lebanon and France3
- Two new species of the bristle millipede genus Pauropsxenus (Diplopoda, Polyxenidae) in mid-Cretaceous Burmese amber2
- Millipeds from Dominican amber, with the description of two new species (Diplopoda: Siphonophoridae) of Siphonophora2
- Die im Bernstein befindlichen Crustaceen, Myriapoden, Arachniden und Apteren der Vorwelt2
- A new early Barremian fossiliferous amber outcrop at Beqaa Kafra (Northern Lebanon)1
- Revision of Phryssonotus burmiticus (Diplopoda, Polyxenida, Synxenidae) in mid-Cretaceous amber from Myanmar1
- Record of pillbug (Armadillidium) and millipede (Polyxenus) remains from the resin lumps of Warkalli Formation (upper Tertiary), Kerala coast1
- Arthropods in Burmese amber1
Each of the 23 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Polyxenida
- Source
- Wikipedia/Commons
- Credit
- Sarefo
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan of Polyxenida was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Polyxenida”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itPenicillata through timeFormations of the Late CretaceousThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Lucas 1840