Araneae
Spiders are air-breathing arthropods that have eight limbs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species diversity among all orders of organisms. Spiders are found worldwide on every continent except Antarctica, and have become established in nearly every land habitat. As of January 2026, 53,680 spider species in 139 families have been recorded by taxonomists. However, there has been debate among scientists about how families should be classified, with over 20 different classifications proposed since 1900.
Michael Gäbler · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- spiderPBDB
- Sits inside
- SerikodiastidaPBDB
- Life habit
- solitaryPBDB, for Arachnida
- Motility
- actively mobilePBDB, for Arachnida
- Environment
- terrestrialPBDB, for Arachnida
Time the record covers
At least 316 million years on record.
The record covers at least 316.9 Mya to 11.7 ka. No occurrence read is dated outside 318.7 Mya to 0 Mya.
2,828 occurrences of Araneae on record, most of them in the Eocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
2,828 occurrences- Carboniferousunder 1%
- Permianunder 1%
- Middle Triassicunder 1%
- Late Triassicunder 1%
- Early Jurassicunder 1%
- Middle Jurassicunder 1%
- Late Jurassicunder 1%
- Early Cretaceous3%
- Late Cretaceous19%
- Paleoceneunder 1%
- Eocene58%
- Oligocene1%
- Miocene14%
- Plioceneunder 1%
- Pleistoceneunder 1%
- Holocene2%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
66 formations
- Florissant Formation53
- Shanwang Formation23
- Niveau du gypse d'Aix Formation20
- Daohugou Formation12
- Raritan Formation · Potomac Group12
- Upper Freshwater-Molasse Formation11
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The named rock units Araneae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 42 further formations hold fewer occurrences each and are not listed. 2,535 occurrences have no formation on record, which is common for older literature.
Araneae and its relatives
EoceneWhere Araneae and its relatives were recorded in the Eocene.
Found in the same rock
7,541 taxa- Ctenobethylus goepperti13 sites
- Gesomyrmex hoernesi13 sites
- Prenolepis henschei13 sites
- Yantaromyrmex geinitzi13 sites
- Archaea paradoxa12 sites
- Balticoroma serafinorum12 sites
- Lasius schiefferdeckeri12 sites
- Prosphyracephala succini12 sites
- Nylanderia pygmaea11 sites
- Tetraponera simplex11 sites
- Coleoptera10 sites
- Formica flori10 sites
- Gorgopsina frenata10 sites
- Orchestina baltica10 sites
- Pachycondyla succinea10 sites
- Plagiolepis klinsmanni10 sites
- Adorator hispidus9 sites
- Bradoponera meieri9 sites
- Diptera9 sites
- Episinus longimanus9 sites
- Formicidae9 sites
- Gnamptogenys europaea9 sites
- Pseudoteutana stigmatosa9 sites
- Segestria9 sites
The Paleobiology Database records at least 261 sites for Araneae, and this reads 240 of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
302 references- Die fossilen Spinnen im dominikanischen Bernstein186
- On extant and fossil (Eocene) European comb-footed spiders (Araneae: Theridiidae), with notes on their subfamilies, and with descriptions of new taxa174
- Footnotes and taxonomic names, in Die im Bernstein befindlichen Crustaceen, Myriapoden, Arachniden und Apteren der Vorwelt118
- On the evolution and the classification of spiders, the Mesozoic spider faunas, and descriptions of new Cretaceous taxa mainly in amber from Myanmar (Burma) (Arachnida: Araneae)97
- Die im Bernstein befindlichen Crustaceen, Myriapoden, Arachniden und Apteren der Vorwelt97
- New and rare fossil spiders (Araneae) in mid Cretaceous amber from Myanmar (Burma), including the description of new extinct families of the suborders Mesothelae and Opisthothelae, as well as notes on the taxonomy, the evolution and the biogeography of the Mesothelae88
- A study of amber spiders88
- Amber Spiders in European Collections84
- Descriptions of new fossil spiders (Arachnida: Araneae) in Late (Mid) Cretaceous Burmese amber with focus on the superfamilies Palpimanoidea and Deinopoidea and members of the RTA-clade, as well as remarks on palaeobehaviour, palaeofauna, taxonomy and phylogenetics70
- The fossil spiders of the family Anapidae s. l. (Araneae) in Baltic, Dominican and Mexican amber and their extant relatives, with the description of a new subfamily Comarominae68
- Fossil spiders (Araneae) of the superfamily Dysderoidea in Baltic and Dominican amber, with revised family diagnoses56
- Fossil spiders of the family Dictynidae s. l., including Cryphoecinae and Hahniinae in Baltic and Dominican amber and copal from Madagascar, and on selected extant Holarctic taxa, with new descriptions and diagnoses54
Each of the 2,828 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 290 further references are not listed. Each entry links to the Paleobiology Database's own record for the work.
Media and terms
Media content1
Picture of Araneae
- Source
- Wikipedia/Commons
- Credit
- Michael Gäbler
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
No 3D scan of Araneae was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Spider”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itSerikodiastida through timeFormations of the EoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Clerck 1757)