Modulus
Modulus is a genus of small sea snails, marine gastropod molluscs in the family Modulidae.

Manuel CABALLER · CC BY 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- ModulidaePBDB
- Diet
- grazerPBDB, for Modulidae
- Life habit
- epifaunalPBDB, for Modulidae
- Motility
- actively mobilePBDB, for Modulidae
- Vision
- limitedPBDB, for Gastropoda
Time the record covers
At least 27 million years on record.
The record covers at least 27.3 Mya to 129 ka. No occurrence read is dated outside 33.9 Mya to 11.7 ka.
95 occurrences of Modulus on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
95 occurrences- Oligocene3%
- Miocene26%
- Pliocene22%
- Pleistocene50%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
32 formations
- Caloosahatchee Formation · Okeechobee Group10
- Chipola Formation · Alum Bluff Group6
- Mare Formation6
- Fort Thompson Formation5
- Anastasia Formation4
- Castilletes Formation4
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The named rock units Modulus has been recovered from, largest first. Drawn where the rock is now, not where it formed. 8 further formations hold fewer occurrences each and are not listed. 20 occurrences have no formation on record, which is common for older literature.
Modulus and its relatives
PleistoceneWhere Modulus and its relatives were recorded in the Pleistocene.
Found in the same rock
2,197 taxa- Cerithium19 sites
- Anadara15 sites
- Bulla striata13 sites
- Olivella13 sites
- Atilia12 sites
- Crepidula plana12 sites
- Turritella12 sites
- Chama11 sites
- Marginella11 sites
- Trachycardium11 sites
- Anomalocardia9 sites
- Arcopsis adamsi9 sites
- Calliostoma9 sites
- Calyptraea centralis9 sites
- Cancellaria9 sites
- Chione cancellata9 sites
- Chlamys9 sites
- Cythara (Mangelia)9 sites
- Epitonium9 sites
- Lamarcka imbricata9 sites
- Nassarius9 sites
- Nuculana acuta9 sites
- Tellina9 sites
- Acteocina8 sites
The Paleobiology Database records 36 sites for Modulus, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
45 references- Unpublished census data from Atlantic coastal plain and circum-Caribbean Neogene assemblages and taxonomic opinions10
- Neogene molluscs, shallow marine paleoenvironments, and chronostratigraphy of the Guajira Peninsula, Colombia7
- Unpublished notes on Caribbean collection of Jack and Winifred Gibson-Smith, Naturhistorisches Museum Basel7
- The Molluscan Fauna of the Alum Bluff Group of Florida6
- Petuch and Roberts, 20074
- Neogene Biostratigraphy of the Charlotte Harbor Area in Southwestern Florida4
- Skwarko, 19943
- The benthic mollusk faunas of two contrasting reef paleosubenvironments: Falmouth Formation (late Pleistocene, last interglacial), Jamaica3
- Distribution of Bivalves and Gastropod in the Pleistocene Ironshore Formation, Grand Cayman, British West Indies3
- Fossil mollusks of Oahu, Hawaii Islands3
- Pliocene Mollusca of Southern Florida3
- Mollusca: Fort Thompson Formation (Late Pleistocene)2
Each of the 95 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 33 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Modulus
- Source
- Wikipedia/Commons
- Credit
- Manuel CABALLER
- Licence
- CC BY 4.0
- Commercial use
- the licence allows it
No 3D scan was found. Smithsonian Open Access, MorphoSource and Sketchfab were all searched.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itModulidae through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Gray 1842