Macrobaenidae
Macrobaenidae is an extinct family of turtles, known from the Early Cretaceous to Paleogene of Laurasia. Their relationships to other turtles and whether they form a monophlyletic group are controversial. They are typically interpreted as stem or crown group cryptodires, but some more recent analyses have found them to lie outside crown group Testudines. Macrobaenids can be distinguished from other testudinatans by the presence of a carotid fenestra, cruciform plastron with strap-like epiplastra, and a lack of extragulars.

Internet Archive Book Images · No restrictions · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- MacrobaenoideaPBDB
- Diet
- omnivorePBDB, for Cryptodira
- Life habit
- aquaticPBDB, for Cryptodira
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- freshwaterPBDB, for Cryptodira
Time the record covers
At least 87 million years on record.
The record covers at least 143.1 Mya to 56 Mya. No occurrence read is dated outside 161.5 Mya to 33.9 Mya.
82 occurrences of Macrobaenidae on record, most of them in the Late Cretaceous. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
82 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
36 formations
- Hell Creek Formation · Montana Group7
- Khodzhakul Formation6
- Hornerstown Formation · Rancocas Group6
- Minhe Formation5
- Lance Formation3
- Severn Formation2
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The named rock units Macrobaenidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 12 further formations hold fewer occurrences each and are not listed. 10 occurrences have no formation on record, which is common for older literature.
Macrobaenidae and its relatives
Late CretaceousWhere Macrobaenidae and its relatives were recorded in the Late Cretaceous.
Found in the same rock
788 taxa- Hadrosauridae15 sites
- Osteopygis backmani13 sites
- Osteopygis emarginatus13 sites
- Testudines12 sites
- Trionychidae12 sites
- Theropoda10 sites
- Baenidae9 sites
- Dromaeosauridae9 sites
- Champsosaurus8 sites
- Chelydridae8 sites
- Adocus7 sites
- Lepisosteidae7 sites
- Osteichthyes7 sites
- Basilemys6 sites
- Ceratopsidae6 sites
- Compsemys victa6 sites
- Crocodylia6 sites
- Trionyx6 sites
- Agomphus pectoralis5 sites
- Anatolemys maximus5 sites
- Aves5 sites
- Euclastes wielandi5 sites
- Hypsilophodontidae5 sites
- Kirgizemys5 sites
The Paleobiology Database records 70 sites for Macrobaenidae, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
47 references- Data on late Mesozoic turtles from the USSR6
- Patterns of geographic variation in latest Cretaceous vertebrates: evidence from the turtle component5
- VI. Vertebrate Palaeontology 6. Fossil reptiles from Mongolia and Kansu5
- Soft-shelled turtles (Trionychidae) from the Cenomanian of Uzbekistan4
- Non-trionychid turtles from the Paleocene of Alberta, Canada3
- Fossil Vertebrate Collections3
- The fossil turtles of North America3
- Synopsis of the Extinct Batrachia, Reptilia and Aves of North America. Part II3
- A large non-marine turtle from the Upper Cretaceous of Alabama and a review of North American “macrobaenids”2
- A European Cenozoic ‘Macrobaenid’: New Data about the Paleocene Arrival of Several Turtle Lineages to Europe2
- Paleogene chelonians from Maryland and Virginia2
- A new eucryptodiran turtle from the Late Cretaceous of North America (Dinosaur Provincial Park, Alberta, Canada)2
Each of the 82 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 35 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Macrobaenidae
- Source
- Wikipedia/Commons
- Credit
- Internet Archive Book Images
- Licence
- No restrictions
- Commercial use
- cannot be read from the licence given — check the record
No 3D scan of Macrobaenidae was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itMacrobaenoidea through timeFormations of the Late CretaceousThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Sukhanov 1964)