Clypeasteroida
Sand dollars are species of flat, burrowing sea urchins belonging to the order Clypeasteroida. Some species within the order, not quite as flat, are known as sea biscuits. Sand dollars can also be called "sand cakes" or "cake urchins".

Frédéric Ducarme · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- sand dollarPBDB
- Sits inside
- NeognathostomataPBDB
- Diet
- detritivorePBDB
- Life habit
- semi-infaunalPBDB
- Motility
- slow-movingPBDB
- Vision
- limitedPBDB, for Echinoidea
- Environment
- marinePBDB, for Echinodermata
Time the record covers
At least 89 million years on record.
The record covers at least 89.8 Mya to 11.7 ka. No occurrence read is dated outside 100.5 Mya to 0 Mya.
1,295 occurrences of Clypeasteroida on record, most of them in the Miocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
1,295 occurrences- Late Cretaceous5%
- Paleoceneunder 1%
- Eocene22%
- Oligocene8%
- Miocene39%
- Pliocene14%
- Pleistocene11%
- Holoceneunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
228 formations
- Vaqueros Formation67
- Tortonian Limestone Formation48
- Castle Hayne Formation · Jackson Group42
- Temblor Formation37
- Moodys Branch Formation · Jackson Group36
- Crystal River Limestone Formation · Ocala Group33
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The named rock units Clypeasteroida has been recovered from, largest first. Drawn where the rock is now, not where it formed. 204 further formations hold fewer occurrences each and are not listed. 285 occurrences have no formation on record, which is common for older literature.
Clypeasteroida and its relatives
MioceneWhere Clypeasteroida and its relatives were recorded in the Miocene.
Found in the same rock
2,084 taxa- Periarchus lyelli51 sites
- Chlamys49 sites
- Ostrea41 sites
- Turritella31 sites
- Clypeaster25 sites
- Cidaris pratti22 sites
- Echinolampas20 sites
- Conus17 sites
- Venericardia17 sites
- Vertebrata17 sites
- Protoscutella conradi16 sites
- Tellina16 sites
- Chama15 sites
- Chlamys deshayesii15 sites
- Flabellum15 sites
- Gigantostrea trigonalis15 sites
- Pacipecten andersoni15 sites
- Vaquerosella merriami15 sites
- Echinocyamus14 sites
- Eburneopecten scintillatus13 sites
- Echinolampas appendiculata13 sites
- Strombus13 sites
- Dentalium12 sites
- Linthia hanoverensis12 sites
The Paleobiology Database records 837 sites for Clypeasteroida, and this reads 240 of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
266 references- Stratigraphic Distribution of Paleocene and Eocene Fossils in the Eastern Gulf Coast Region74
- The Vaqueros Formation, Lower Miocene of California. I. Paleontology62
- The West American Cenozoic Echinoidea52
- Palaeoenvironmental reconstruction based on non-geniculate coralline red algal assemblages in Miocene limestone of central Crete48
- The Echinoids of the Middle Eocene Warley Hill Formation, Santee Limestone, and Castle Hayne Limestone of North and South Carolina39
- Diversity patterns and biostratigraphy of Cenozoic echinodems from Florida37
- Geology and paleontology of the Santa Maria District California35
- Stratigraphy and paleontology of the marine Neogene formations of the Coalinga region, California32
- The Mesozoic and Cenozoic Echinodermata of the United States29
- Cenozoic irregular echinoids of the Eastern United States28
- Los Equiodermos fosiles de Cuba28
- Late Cretaceous (Maastrichtian) echinoid-mollusc-coral assemblages and palaeoenvironments from a Tethyan carbonate platform succession, northern Oman Mountains23
Each of the 1,295 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 254 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content2
Picture of Clypeasteroida
- Source
- Wikipedia/Commons
- Credit
- Frédéric Ducarme
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Sand dollar”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itNeognathostomata through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Agassiz 1836