Cassidinae
The Cassidinae are a subfamily of the leaf beetles, or Chrysomelidae. The antennae arise close to each other and some members have the pronotal and elytral edges extended to the side and covering the legs so as to give them the common name of tortoise beetles. Some members, such as in the tribe Hispini, are notable for the spiny outgrowths to the pronotum and elytra.

No machine-readable author provided. Shyamal assumed (based on copyright claims). · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- tortoise beetlePBDB
- Sits inside
- ChrysomelidaePBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 45 million years on record.
The record covers at least 48.07 Mya to 2.58 Mya. No occurrence read is dated outside 56 Mya to 1.8 Mya.
42 occurrences of Cassidinae on record, most of them in the Eocene. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
42 occurrences- Eocene50%
- Oligocene14%
- Miocene33%
- Plioceneunder 1%
- Pleistocene2%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
7 formations
- Upper Freshwater-Molasse Formation6
- Niveau du gypse d'Aix Formation4
- Eifel Formation2
- Green River Formation2
- Florissant Formation1
- Shanwang Formation1
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The named rock units Cassidinae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 25 occurrences have no formation on record, which is common for older literature.
Cassidinae and its relatives
EoceneWhere Cassidinae and its relatives were recorded in the Eocene.
Found in the same rock
2,021 taxa- Cassida blancheti5 sites
- Curculionidae5 sites
- Cerambycidae4 sites
- Libellulidae4 sites
- Zygoptera4 sites
- Apis3 sites
- Bibio curtisii3 sites
- Buprestis3 sites
- Cassida3 sites
- Coleoptera3 sites
- Entiminae3 sites
- Entimini3 sites
- Gonophora bremii3 sites
- Plagiodera lyelliana3 sites
- Plecia bucklandi3 sites
- Plecia foersteri3 sites
- Plecia livida3 sites
- Protainophthalmus asperulus3 sites
- Protainophthalmus margarus3 sites
- Termitidae3 sites
- Agrilini2 sites
- Anisoptera2 sites
- Anthaxia haupti2 sites
- Anthaxia viridicyanea2 sites
The Paleobiology Database records 31 sites for Cassidinae, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
25 references- Beitrag zur Kenntnis der eozänen Arthopodenfauna des Geiseltales4
- Heer, 18474
- Die Käfer (Coleoptera) aus der eozänen Braunkohle des Geiseltales3
- Ueber die fossilen Insekten von Aix in der Provence3
- Eocene tortoise beetles from the Green River Formation in Colorado, U.S.A. (Coleoptera: Chrysomelidae: Cassidinae)2
- Systematik, Phylogenie, Taphonomie und Paläoökologie der Insekten aus dem Mittel-Eozän des Eckfelder Maares, Vulkaneifel2
- Dominican and Mexican amber chrysomelids, with descriptions of two new species2
- Deux Dicladispa fossiles des gisements de diatomites Néogènes de Bes-Konak (Turquie) et de Murat (France) (Coleoptera, Chrysomelidae, Hispinae)2
- Hispinen aus baltischen Bernstein2
- Die eozäne Insektenfauna des Geiseltales2
- Recherches sur les insectes fossiles des terrains Tertiaires de la France, deuxième partie, insectes fossiles d'Aix en Provence2
- A new Sceloenopla (Coleoptera, Chrysomelidae, Hispinae) from Dominican amber1
Each of the 42 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 13 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content2
Picture of Cassidinae
- Source
- Wikipedia/Commons
- Credit
- No machine-readable author provided. Shyamal assumed (based on copyright claims).
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
3D model
- Source
- Sketchfab
- Credit
- Yale Peabody Museum
- Licence
- CC Attribution
- Commercial use
- cannot be read from the licence given — check the record
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Cassidinae”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itChrysomelidae through timeFormations of the EoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Gyllenhal 1813