Bothremydidae
Bothremydidae is an extinct family of side-necked turtles (Pleurodira) known from the Cretaceous and Cenozoic. They are closely related to Podocnemididae, and are amongst the most widely distributed pleurodire groups, with their fossils having been found in Africa, India, the Middle East, Europe, North America and South America. Bothremydids were aquatic turtles with a high morphological diversity, indicative of generalist, molluscivorous, piscivorous and possibly herbivorous grazing diets, with some probably capable of suction feeding. Unlike modern pleurodires, which are exclusively freshwater, bothremydids inhabited freshwater, marine and coastal environments. Their marine habits allowed bothremydids to disperse across oceanic barriers into Europe and North America during the early Late Cretaceous (Cenomanian). The youngest records of the group are indeterminate remains from Saudi Arabia and Oman, dating to the Miocene.

Ghedo · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- PleurodiraPBDB
- Diet
- omnivorePBDB, for Pleurodira
- Life habit
- amphibiousPBDB, for Pleurodira
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- freshwaterPBDB, for Pleurodira
Time the record covers
At least 80 million years on record.
The record covers at least 121.4 Mya to 41.03 Mya. No occurrence read is dated outside 125.77 Mya to 33.9 Mya.
246 occurrences of Bothremydidae on record, most of them in the Late Cretaceous. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
246 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
72 formations
- Conques Formation · Tremp Group17
- Hornerstown Formation · Rancocas Group13
- Tar Heel Formation · Black Creek Group11
- Landana Formation10
- Aguja Formation10
- Dakhla Formation7
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The named rock units Bothremydidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 48 further formations hold fewer occurrences each and are not listed. 60 occurrences have no formation on record, which is common for older literature.
Bothremydidae and its relatives
Late CretaceousWhere Bothremydidae and its relatives were recorded in the Late Cretaceous.
Found in the same rock
1,026 taxa- Hadrosauridae25 sites
- Eusuchia21 sites
- Dromaeosauridae20 sites
- Lepisosteidae19 sites
- Squalicorax pristodontus19 sites
- Theropoda19 sites
- Dinosauria18 sites
- Elasmosauridae15 sites
- Chelonioidea14 sites
- Osteichthyes14 sites
- Anura13 sites
- Cretalamna appendiculata13 sites
- Nodosauridae13 sites
- Teleostei13 sites
- Polysternon12 sites
- Crocodylia11 sites
- Mosasauridae11 sites
- Pycnodontidae11 sites
- Squalicorax kaupi11 sites
- Taphrosphyini11 sites
- Cretalamna maroccana10 sites
- Crocodyliformes10 sites
- Enchodus elegans10 sites
- Plioplatecarpinae10 sites
The Paleobiology Database records 194 sites for Bothremydidae, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
109 references- Evolution of the side-necked turtles: The families Bothremydidae, Euraxemydidae, and Araripemydidae42
- Chronostratigraphy and new vertebrate sites from the upper Maastrichtian of Huesca (Spain), and their relation with the K/Pg boundary14
- New material of North American side-necked turtles (Pleurodira: Bothremydidae)14
- Shell anatomy of the African Paleocene bothremydid turtle Taphrosphys congolensis and systematic implications within Taphrosphyini9
- Turtles of the Aguja and Javelina Formations, Upper Cretaceous (Campanian – Maastrichtian), West Texas7
- Marine vertebrate faunas from the Maastrichtian phosphates of Benguérir (Ganntour Basin, Morocco): Biostratigraphy, palaeobiogeography and palaeoecology5
- Kinkonychelys, a new side-necked turtle (Pelomedusoides: Bothremydidae) from the Late Cretaceous of Madagascar5
- Descripción de los restos de quelonios del Maastrichtiense superior de Aren y Serraduy (Huesca) [Description of chelonian remains from the Upper Maastrichtian of Aren and Seraduy (Huesca)]5
- Synopsis of the Extinct Batrachia, Reptilia and Aves of North America. Part II5
- New bothremydid turtle (Testudines, Pleurodira) from the Paleocene of Northeastern Colombia4
- New dinosaur sites correlated with Upper Maastrichtian pelagic deposits in the Spanish Pyrenees: implications for the dinosaur extinction pattern in Europe4
- The Cretaceous/Tertiary mass extinction event in the North Atlantic coastal plain4
Each of the 246 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 97 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Bothremydidae
- Source
- Wikipedia/Commons
- Credit
- Ghedo
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan of Bothremydidae was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itPleurodira through timeFormations of the Late CretaceousThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Baur 1891)