Testudinidae
Tortoises are reptiles of the family Testudinidae of the order Testudines. Like other testudines, tortoises have a shell to protect from predation and other threats. The shell in tortoises is generally hard, and like other members of the suborder Cryptodira, they retract their necks and heads directly backward into the shell to protect them.

Childzy at en.wikipedia · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- TestudinoideaPBDB
- Diet
- herbivore, herbivorePBDB
- Life habit
- ground dwellingPBDB
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB
Time the record covers
At least 100 million years on record.
The record covers at least 100.5 Mya to 11.7 ka. No occurrence read is dated outside 113.2 Mya to 0 Mya.
1,488 occurrences of Testudinidae on record, most of them in the Miocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
1,488 occurrences- Early Cretaceousunder 1%
- Late Cretaceousunder 1%
- Paleocene1%
- Eocene8%
- Oligocene6%
- Miocene43%
- Pliocene12%
- Pleistocene26%
- Holocene4%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
255 formations
- Collon Cura Formation31
- Huayquerías Formation22
- Koobi Fora Formation21
- Brule Formation · White River Group19
- Vogel River Series Formation16
- Upper Freshwater Molasse Formation · Molasse Group13
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The named rock units Testudinidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 231 further formations hold fewer occurrences each and are not listed. 863 occurrences have no formation on record, which is common for older literature.
Testudinidae and its relatives
MioceneWhere Testudinidae and its relatives were recorded in the Miocene.
Found in the same rock
3,776 taxa- Geochelone67 sites
- Testudo51 sites
- Trionyx39 sites
- Equus36 sites
- Hipparion34 sites
- Bovidae33 sites
- Gazella30 sites
- Rana29 sites
- Crocodylus27 sites
- Aves26 sites
- Ceratotherium26 sites
- Rhinocerotidae26 sites
- Crocodylus niloticus25 sites
- Euthecodon brumpti25 sites
- Trionychidae25 sites
- Polypterus24 sites
- Clarias23 sites
- Hystrix21 sites
- Ophisaurus21 sites
- Lates20 sites
- Osteichthyes20 sites
- Protopterus20 sites
- Canis19 sites
- Cercopithecidae19 sites
The Paleobiology Database records 1,250 sites for Testudinidae, and this reads 240 of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
642 references- Contribution a l'etude des Cheloniens. Cheloniens continentaux due Cretace et du Tertiare de France71
- Testudinid turtle remains from the Late Miocene palaeo-island of Gargano, Italy, and an overview of Mediterranean insular tortoises30
- Tortoises and turtles from Huayquerías del Este (late Miocene–early Pliocene), Mendoza, Argentina29
- Morphology of the mid-sized tortoises (Testudines: Testudinidae) from the Middle Miocene of Northwestern Chubut27
- A fossil and zooarchaeological history of the gopher tortoise (Gopherus polyphemus) in the Southeastern United States23
- A revision of Anhuichelys Yeh, 1979, the earliest known stem Testudinidae (Testudines: Cryptodira) from the Paleocene of China16
- The Fossil Chelonians of Greece. Systematics - Evolution - Stratigraphy - Palaeoecology16
- Laetoli: a Pliocene Site in Northern Tanzania16
- New generic proposal for the European Neogene large testudinids (Cryptodira) and the first phylogenetic hypothesis for the medium and large representatives of the European Cenozoic record14
- Neogene Testudines of Germany. Their stratigraphical and ecological evaluation14
- Stratigraphy and mammal faunas of the Brackwassermolasse in the surroundings of Ulm (Southwest Germany)13
- Skeletochronology: a method for determing the individual age and growth of modern and fossil tortoises (Reptilia: Testudines)12
Each of the 1,488 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 630 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Testudinidae
- Source
- Wikipedia/Commons
- Credit
- Childzy at en.wikipedia
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
No 3D scan of Testudinidae was found in Smithsonian Open Access and Sketchfab.
MorphoSource was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Tortoise”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itTestudinoidea through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Batsch 1788