Spisula
Spisula is a genus of medium-sized to large marine bivalve mollusks or clams in the subfamily Mactrinae of the family Mactridae, commonly known as surf clams or trough shells.

Philippe MAESTRATI · CC BY 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- MactrinaePBDB
- Diet
- suspension feederPBDB
- Life habit
- infaunalPBDB
- Motility
- facultatively mobilePBDB
- Vision
- blindPBDB, for Bivalvia
Time the record covers
At least 59 million years on record.
The record covers at least 59.24 Mya to 11.7 ka. No occurrence read is dated outside 66 Mya to 0 Mya.
576 occurrences of Spisula on record, most of them in the Eocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
576 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
134 formations
- Moodys Branch Formation · Jackson Group45
- Choptank Formation · Chesapeake Group20
- Shoal River Formation · Alum Bluff Group17
- Norwich Crag Formation · Crag Group16
- Gosport Sand Formation · Claiborne Group15
- Oak Grove Formation · Alum Bluff Group15
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The named rock units Spisula has been recovered from, largest first. Drawn where the rock is now, not where it formed. 110 further formations hold fewer occurrences each and are not listed. 93 occurrences have no formation on record, which is common for older literature.
Spisula and its relatives
EoceneWhere Spisula and its relatives were recorded in the Eocene.
Found in the same rock
3,025 taxa- Turritella42 sites
- Gastropoda38 sites
- Eburneopecten scintillatus37 sites
- Bivalvia34 sites
- Calyptraea (Trochita) aperta29 sites
- Tellinidae29 sites
- Caestocorbula wailesiana28 sites
- Venericardia (Rotundicardia) diversidentata28 sites
- Cadulus27 sites
- Callista annexa27 sites
- Corbula densata27 sites
- Myrtaea (Myrtaea) curta26 sites
- Hipponix25 sites
- Nucula (Nucula) sphenopsis25 sites
- Diplodonta (Diplodonta) ungulina23 sites
- Kapalmerella alveata23 sites
- Naticidae23 sites
- Bathytormus flexurus22 sites
- Calorhadia (Litorhadia) mater22 sites
- Actinopterygii21 sites
- Bryozoa21 sites
- Callista21 sites
- Crassinella pygmaea21 sites
- Dentalium21 sites
The Paleobiology Database records 344 sites for Spisula, and this reads 240 of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
135 references- Unpublished census data from Atlantic coastal plain and circum-Caribbean Neogene assemblages and taxonomic opinions41
- Marine Molluscan Assemblages in the Early Pleistocene of Sidestrand, Bramerton and the Royal Society Borehole at Ludham, Norfolk40
- Fossil assemblages of a marine transgressive sand, Moody''s Branch Formation (Upper Eocene), Louisiana and Mississippi27
- The Molluscan Fauna of the Alum Bluff Group of Florida27
- Ecological control of evolutionary rates in Paleocene - Eocene marine molluscs24
- Stratigraphy, invertebrate taphonomy and depositional history of the Miocene Calvert and Choptank Formations, Atlantic Coastal Plain21
- Catalogue of the Paleocene and Eocene mollusca of the southern and eastern United States. Part 1. Pelecypoda, Amphineura, Pteropoda, Scaphopoda and Cephalopoda20
- Studies on the marine Pleistocene: Part II. The marine Pleistocene mollusks of eastern North America19
- The Neogene Bivalvia (Heterodonta and Anomalodesmata) and Scaphopoda from Kallo and Doel (Oost Vlaanderen, Belgium)15
- Lower Oligocene Bivalvia of the Vicksburg Group in Mississippi15
- Die Fauna des marinen Miozäns von Kevelaer (Niederrhein). Band 2 Bivalvia, Scaphopoda, Cephalopoda, Bryozoa, Annelida, Brachiopoda13
- Stratigraphic Distribution of Paleocene and Eocene Fossils in the Eastern Gulf Coast Region13
Each of the 576 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 123 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content2
Picture of Spisula
- Source
- Wikipedia/Commons
- Credit
- Philippe MAESTRATI
- Licence
- CC BY 4.0
- Commercial use
- the licence allows it
CT scan
- Source
- MorphoSource
- Licence
- not stated by the source
- Commercial use
- cannot be read from the licence given — check the record
- Specimen
- Spisula solidissima · USNM:IZ:USNM 37020
The description on this site
- Source
- Wikipedia, “Spisula”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itMactrinae through timeFormations of the EoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Gray 1837)