Pulmonata
Pulmonata or pulmonates is an informal group of snails and slugs characterized by the ability to breathe air, by virtue of having a pallial lung instead of a gill, or gills. The group includes many land and freshwater families, and several marine families.

Ghost_Slug_adult.jpg : ©Amgueddfa Cymru - National Museum Wales derivative work: Anna Frodesiak ( talk ) · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- pulmonatePBDB
- Sits inside
- GastropodaPBDB
- Life habit
- epifaunalPBDB, for Gastropoda
- Motility
- actively mobilePBDB
- Vision
- limitedPBDB, for Gastropoda
- Environment
- terrestrialPBDB
Time the record covers
At least 330 million years on record.
The record covers at least 330.3 Mya to 11.7 ka. No occurrence read is dated outside 335.9 Mya to 0 Mya.
40 occurrences of Pulmonata on record, most of them in the Miocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
40 occurrences- Carboniferous3%
- Early Cretaceous5%
- Late Cretaceous8%
- Eocene13%
- Oligoceneunder 1%
- Miocene44%
- Pleistocene23%
- Holocene5%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
11 formations
- Griman Creek Formation · Rolling Downs Group3
- Upper Freshwater Molasse Formation · Molasse Group2
- Randeck Maar Formation2
- Willwood Formation1
- Sappa Formation1
- Blesa Formation1
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The named rock units Pulmonata has been recovered from, largest first. Drawn where the rock is now, not where it formed. 25 occurrences have no formation on record, which is common for older literature.
Pulmonata and its relatives
MioceneWhere Pulmonata and its relatives were recorded in the Miocene.
Found in the same rock
1,154 taxa- Discus12 sites
- Discus pleuradrus9 sites
- Gastrocopta acuminata8 sites
- Limax8 sites
- Bufo6 sites
- Gastrocopta nouletiana6 sites
- Gyraulus applanatus6 sites
- Lymnaea dilatata6 sites
- Pseudochloritis incrassata6 sites
- Vertigo6 sites
- Carychium5 sites
- Granaria5 sites
- Gyraulus5 sites
- Leucochroopsis kleinii5 sites
- Pisidium5 sites
- Planorbarius mantelli5 sites
- Sphaerium5 sites
- Apula coarctata4 sites
- Argna oppoliensis4 sites
- Cepaea4 sites
- Hyla4 sites
- Hypsilophodontidae4 sites
- Pomatias4 sites
- Rana4 sites
The Paleobiology Database records 34 sites for Pulmonata, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
28 references- Terrestrial and Freshwater Turtles of Early Cretaceous Australia3
- Depositional dynamics and the record of ecosystem stability: early Eocene faunal gradients in the Pyrenean foreland, Spain3
- Synopsis of the Late Miocene mollusc fauna of the classical sections Richardhof and Eichkogel in the Vienna Basin (Austria, Pannonian, MN 9-MN11)3
- Opole (Poland) - a key locality for middle Miocene terrestrial mollusc faunas2
- Fossil land and freshwater gastropods from the Middle Miocene of Bechingen and Daugendorf, southwestern Germany2
- Fossil gastropods from Miocene Lake Randeck Maar and its hinterland (SW Germany)2
- The fossil pulmonate snails of Sandelzhausen (Early/Middle Miocene, Germany) (Hygrophila, Punctoidea and limacoids)2
- L'environnement côtier des Alpes-Maritimes à la fin de la transgression versilienne d'après l'étude biostratigraphique du site de l'Etoile à Nice (France)2
- Eshelman and Hager, 19842
- Early Miocene continental gastropods from new localities of the Molasse Basin in southern Germany1
- Biostratigraphy and palaeoecology of a Middle Miocene (Karpathian, MN 5) fauna from the northern margin of the North Alpine Foreland Basin (Oggenhausen 2, SW' Germany)1
- Biostratigraphy of Middle Miocene (Sarmatian) wetland systems in an Eastern Alpine intramontane basin (Gratkorn Basin, Austria): the terrestrial gastropod approach1
Each of the 40 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 16 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Pulmonata
- Source
- Wikipedia/Commons
- Credit
- Ghost_Slug_adult.jpg : ©Amgueddfa Cymru - National Museum Wales derivative work: Anna Frodesiak ( talk )
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
No 3D scan of Pulmonata was found in Smithsonian Open Access and Sketchfab.
MorphoSource was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Pulmonata”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itGastropoda through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Cuvier 1817