Pugnax

via iDigBio · CC0 · via iDigBio. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- PugnacidaePBDB
- Diet
- suspension feederPBDB, for Rhynchonellida
- Life habit
- epifaunalPBDB, for Rhynchonellida
- Motility
- stationaryPBDB, for Rhynchonellida
- Vision
- blindPBDB, for Rhynchonellata
- Environment
- marinePBDB, for Lophophorata
Time the record covers
At least 139 million years on record.
The record covers at least 393.47 Mya to 254.14 Mya. No occurrence read is dated outside 419.62 Mya to 251.902 Mya.
59 occurrences of Pugnax on record, most of them in the Carboniferous. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
59 occurrences- Devonian36%
- Carboniferous38%
- Permian27%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
26 formations
- Wargal Formation7
- Weaver Beds Formation5
- Eyam Limestone Formation5
- Glubokaya Formation5
- Waterhouses Limestone Formation2
- Taemas Formation · Murrumbidgee Group2
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The named rock units Pugnax has been recovered from, largest first. Drawn where the rock is now, not where it formed. 2 further formations hold fewer occurrences each and are not listed. 10 occurrences have no formation on record, which is common for older literature.
Pugnax and its relatives
CarboniferousWhere Pugnax and its relatives were recorded in the Carboniferous.
Found in the same rock
935 taxa- Chonetes9 sites
- Brachythyris8 sites
- Camarotoechia8 sites
- Dielasma8 sites
- Schizophoria resupinata8 sites
- Spirifer8 sites
- Betaneospirifer moosakhailensis7 sites
- Dictyoclostus7 sites
- Reticularia7 sites
- Stenoscisma purdoni7 sites
- Athyris6 sites
- Avonia6 sites
- Cleiothyridina subexpansa6 sites
- Eomarginifera6 sites
- Linoproductus6 sites
- Linoproductus cora6 sites
- Martinia glabra6 sites
- Phricodothyris6 sites
- Plicatifera6 sites
- Pustula6 sites
- Terebratuloidea davidsoni6 sites
- Amplexus coralloides5 sites
- Angiospirifer bisulcatus5 sites
- Callispirina ornata5 sites
The Paleobiology Database records 40 sites for Pugnax, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
32 references- The stratigraphy of the Carboniferous rocks of the Weaver Hills District, North Staffordshire8
- Rzhonsnitskaya, 19686
- Brachiopoda and Mollusca from the Productus limestones of the Salt Range6
- The origin and significance of the distribution of shelly macrofauna in Late Dinantian carbonate mud mounds of Derbyshire5
- Unterdevonische Brachiopoden aus dem hellen Crinoidenschutt- und Riffkalk der Seewarte (zentrale Karnische Alpen/Kärnten/Österreich)2
- Famennian faunal zones in western Canada2
- The brachiopods and the boundary of Late Carboniferous-Early Permian in Longlin region, Guangxi2
- Brachiopods of the Murrumbidgee Group, Taemas, New South Wales2
- A detailed account of the fauna and age of a Waulsortian Reef Knoll Limestone and associated shales, Feltrim, Co., Dublin2
- Productus Limestone Fossils, Part IV, fas. 22
- Frasnian-Famennian extinction and recovery of rhynchonellid brachiopods from the East European Platform1
- Wenlockian to Emsian communities of the Carnic Alps (Austria and Italy)1
Each of the 59 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 20 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content3

Picture of Pugnax
- Source
- iDigBio
- Credit
- via iDigBio
- Licence
- CC0
- Commercial use
- the licence allows it
CT scan
- Source
- MorphoSource
- Licence
- In Copyright
- Commercial use
- cannot be read from the licence given — check the record
- Specimen
- Betta pugnax · ummz:fish:209908
3D model
- Source
- Sketchfab
- Licence
- CC0 Public Domain
- Commercial use
- the licence allows it
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itPugnacidae through timeFormations of the CarboniferousThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0.