Protungulatum
Protungulatum is an extinct genus of eutherian mammals within extinct family Protungulatidae, and is possibly one of the earliest known placental mammals in the fossil record, that lived in North America from the Late Cretaceous to early Paleocene.

Uncredited · CC0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- ArctocyonidaePBDB
- Diet
- omnivorePBDB
- Life habit
- ground dwellingPBDB, for Arctocyonidae
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Eutheria
Time the record covers
At most 8.8 million years on record.
Every occurrence read is dated to somewhere between 72.2 Mya and 63.4 Mya, and nothing in the dating separates them, so that window is what the record shows rather than a span.
64 occurrences of Protungulatum on record, most of them in the Paleocene. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
64 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
10 formations
- Hell Creek Formation41
- Ferris Formation8
- Fort Union Formation7
- Lance Formation1
- Frenchman Formation1
- Ravenscrag Formation1
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The named rock units Protungulatum has been recovered from, largest first. Drawn where the rock is now, not where it formed. 1 occurrence has no formation on record, which is common for older literature.
Protungulatum and its relatives
PaleoceneWhere Protungulatum and its relatives were recorded in the Paleocene.
Found in the same rock
218 taxa- Mesodma24 sites
- Stygimys kuszmauli24 sites
- Baioconodon nordicus18 sites
- Oxyprimus erikseni18 sites
- Mimatuta17 sites
- Procerberus formicarum17 sites
- Cimexomys minor16 sites
- Mimatuta morgoth16 sites
- Stygimys gratus16 sites
- Mesodma thompsoni8 sites
- Mimatuta minuial8 sites
- Valenopsalis joyneri8 sites
- Mesodma formosa7 sites
- Mesodma hensleighi6 sites
- Trionychidae6 sites
- Mesodma ambigua5 sites
- Oxyprimus5 sites
- Periptychidae5 sites
- Teleostei5 sites
- Thylacodon pusillus5 sites
- Catopsalis alexanderi4 sites
- Champsosaurus4 sites
- Lepisosteidae4 sites
- Thylacodon montanensis4 sites
The Paleobiology Database records 50 sites for Protungulatum, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
19 references- The Bug Creek problem and the Cretaceous-Tertiary transition at McGuire Creek, Montana31
- A new important record of earliest Cenozoic mammalian history: geologic setting, Multituberculata, and Peradectia8
- Mammal evolution near the Cretaceous-Tertiary Boundary4
- The beginning of the Age of Mammals3
- Early mammalian recovery after the end-Cretaceous mass extinction: A high-resolution view from McGuire Creek area, Montana, USA2
- Gradual dinosaur extinction and simultaneous ungulate radiation in the Hell Creek Formation2
- A study of Mammalia and geology across the Cretaceous-Tertiary boundary in Garfield County, Montana2
- New mammalian local faunas from the first ca. 80 ka of the Paleocene in northeastern Montana and a revised model of biotic recovery from the Cretaceous–Paleogene mass extinction1
- Earliest Palaeocene purgatoriids and the initial radiation of stem primates1
- A new earliest Paleocene (Puercan) mammalian fauna from Colorado's Denver Basin, U.S.A1
- A new earliest Paleocene (Puercan) arctocyonid mammal from the Fort Union Formation, Great Divide Basin, Wyoming, and its phylogenetic position among early condylarths1
- Protungulatum, confirmed Cretaceous occurrence of an otherwise Paleocene eutherian (placental?) mammal1
Each of the 64 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 7 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Protungulatum
- Source
- Wikipedia/Commons
- Credit
- nobody named on the record
- Licence
- CC0
- Commercial use
- the licence allows it
No 3D scan of Protungulatum was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itArctocyonidae through timeFormations of the PaleoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Sloan and Van Valen 1965