Protictitherium
Protictitherium is an extinct genus of hyena that lived across Europe and Asia during the Middle and Late Miocene, it is often considered to be the first hyena since it contains some of the oldest fossils of the family. They were especially prolific in Turkey, where every species has been registered.

PePeEfe · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- IctitheriinaePBDB
- Diet
- carnivorePBDB, for Hyaenidae
- Life habit
- ground dwellingPBDB, for Hyaenidae
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Eutheria
Time the record covers
At least 7.2 million years on record.
The record covers at least 15.98 Mya to 8.725 Mya. No occurrence read is dated outside 20.45 Mya to 5.09 Mya.
22 occurrences of Protictitherium on record, most of them in the Miocene. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
22 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
7 formations
- Nea Messimvria Formation3
- Ville Formation1
- Brackwater Molasse Formation · Molasse Group1
- Chersonian Formation · Balta Group1
- Dytiko Formation1
- Detritic-Carbonated Unit of Subirats Formation1
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The named rock units Protictitherium has been recovered from, largest first. Drawn where the rock is now, not where it formed. 13 occurrences have no formation on record, which is common for older literature.
Protictitherium and its relatives
MioceneWhere Protictitherium and its relatives were recorded in the Miocene.
Found in the same rock
518 taxa- Gomphotherium angustidens7 sites
- Brachypotherium brachypus6 sites
- Heteroxerus6 sites
- Lagopsis6 sites
- Spermophilinus bredai6 sites
- Anchitherium aurelianense5 sites
- Aves5 sites
- Bunolistriodon5 sites
- Copemys5 sites
- Dorcatherium crassum5 sites
- Procervulus dichotomus5 sites
- Prolagus oeningensis5 sites
- Soricidae5 sites
- Steneofiber depereti5 sites
- Testudinidae5 sites
- Testudo5 sites
- Amphicyon giganteus4 sites
- Amphicyonidae4 sites
- Amphimoschus ponteleviensis4 sites
- Anchitherium4 sites
- Dicrocerus elegans4 sites
- Dorcatherium guntianum4 sites
- Dorcatherium naui4 sites
- Gazella4 sites
The Paleobiology Database records 18 sites for Protictitherium, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
17 references- New Miocene locality in Turkey with evidence of the origin of Ramapithecus and Sivapithecus3
- A new protictithere from the late Miocene hominoid locality Ravin de la Pluie of Axios Valley (Macedonia, Greece)2
- Le gisement a mammiferes miocenes de Castelnau-d'Arbieu (Gers). Donnees nouvelles et implications biostratigraphiques2
- Prosansanosmilus peregrinus, ein neuer machairodontider Felide aus dem Miocän Deutschlands und Frankreichs2
- Kayaca, a new vertebrate locality from the Upper Miocene of Türkiye and its importance for the Turolian biogeography of the eastern peri-Mediterranean region1
- Els Casots (Subirats, Catalonia), a key site for the Miocene vertebrate record of Southwestern Europe1
- Le site paléontologique du Grand Morier (Pont-Boutard, Indre-et-Loire, France): contexte géologique et détail biostratigraphique des formations cénozoïques à partir des assemblages de vertébrés fossiles1
- Late Miocene large mammals from Mahmutgazi, Denizli province, Western Turkey1
- Basal middle Miocene Listriodontinae (Suidae, Artiodactyla) from Madrid, Spain1
- Late Miocene bony fishes from Pocsesti (Republic of Moldova)1
- Los yacimientos de vertebrados continentales del Aragoniense superior (Mioceno medio) de Toril, Cuenca de Calatayud-Daroca1
- Les vertébrés du Miocène moyen de Contres (Loir-et-Cher, France)1
Each of the 22 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 5 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Protictitherium
- Source
- Wikipedia/Commons
- Credit
- PePeEfe
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan was found. Smithsonian Open Access, MorphoSource and Sketchfab were all searched.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itIctitheriinae through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Kretzoi 1938