Palaeognathae
Palaeognathae is an infraclass of birds, called paleognaths or palaeognaths. It is one of the two infraclasses of birds, the other being Neognathae, both of which form Neornithes. Palaeognathae contains five extant orders consisting of four flightless lineages, termed ratites, and one flying lineage, the Neotropic tinamous. There are 47 species of tinamous, five of kiwis (Apteryx), three of cassowaries (Casuarius), one of emus (Dromaius), two of rheas (Rhea) and two of ostriches (Struthio). Recent research has indicated that paleognaths are monophyletic but the traditional taxonomic split between flightless and flighted forms is incorrect; tinamous are within the ratite radiation, meaning flightlessness arose independently multiple times via parallel evolution.

Bernard DUPONT User:Kore User:Kadellar Frank Vincentz The.Rohit No machine-readable author provided. Manuel Anastácio… · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- NeornithesPBDB
- Diet
- carnivorePBDB, for Coelurosauria
- Life habit
- ground dwelling, solitaryPBDB, for Theropoda
- Motility
- actively mobilePBDB, for Theropoda
- Environment
- terrestrialPBDB, for Theropoda
Time the record covers
At least 65 million years on record.
The record covers at least 66 Mya to 11.7 ka. No occurrence read is dated outside 72.2 Mya to 0 Mya.
389 occurrences of Palaeognathae on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
389 occurrences- Late Cretaceousunder 1%
- Paleocene3%
- Eocene6%
- Oligocene1%
- Miocene18%
- Pliocene12%
- Pleistocene31%
- Holocene29%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
75 formations
- Olduvai Formation · Vogel River Series Group14
- Hillgrove Formation12
- Monte Hermoso Formation10
- Santa Cruz Formation9
- Bannockburn Formation · Manuherikia Group7
- London Clay Formation · Thames Group6
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The named rock units Palaeognathae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 51 further formations hold fewer occurrences each and are not listed. 226 occurrences have no formation on record, which is common for older literature.
Palaeognathae and its relatives
PleistoceneWhere Palaeognathae and its relatives were recorded in the Pleistocene.
Found in the same rock
3,025 taxa- Struthio48 sites
- Gazella35 sites
- Aves31 sites
- Hipparion28 sites
- Bovidae24 sites
- Mammalia24 sites
- Reptilia24 sites
- Equus23 sites
- Struthio camelus23 sites
- Canis19 sites
- Gallirallus australis19 sites
- Pachyornis elephantopus18 sites
- Alcelaphini17 sites
- Amphibia17 sites
- Clarias16 sites
- Osteichthyes16 sites
- Dinornithiformes15 sites
- Cercopithecidae14 sites
- Emeus crassus14 sites
- Suidae14 sites
- Testudines14 sites
- Viverridae14 sites
- Chenonetta finschi13 sites
- Coturnix novaezelandiae13 sites
The Paleobiology Database records 260 sites for Palaeognathae, and this reads 240 of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
173 references- Quaternary fossil fauna of South Canterbury, South Island, New Zealand56
- Quaternary fossil faunas from caves in Takaka Valley and on Takaka Hill, northwest Nelson, South Island, New Zealand27
- Late-Pleistocene avifaunas from Cape Wanbrow, Otago, South Island, New Zealand21
- A remarkable fossil and archaeological avifauna from Marfells Beach, Lake Grassmere, South Island, New Zealand14
- The fossil history of the emus, Dromaius (Aves: Dromaiinae)9
- Catalogue of Tertiary Fossil Sites of the Land Mammals in the U.S.S.R.9
- Late Miocene continental birds from the Cerro Azul Formation in the Pampean region (central- southern Argentina)7
- Lothagam: the dawn of humanity in eastern Africa7
- Late Cenozoic vertebrates from the Southern Pampean region: systematic and bio-chronostratigraphic update6
- Late Cenozoic mammal bio-chronostratigraphy in southwestern Buenos Aires Province, Argentina6
- Holocene bird subfossils from the sites of Ampasambazimba, Antsirabe and Ampoza, Madagascar: changes in the avifauna of south central Madagascar over the past few millennia6
- Birds of the British Lower Eocene6
Each of the 389 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 161 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Palaeognathae
- Source
- Wikipedia/Commons
- Credit
- Bernard DUPONT User:Kore User:Kadellar Frank Vincentz The.Rohit No machine-readable author provided. Manuel Anastácio…
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan of Palaeognathae was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itNeornithes through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Pycraft 1900)