Monstersauria
Monstersauria is a clade of anguimorph lizards, defined as all taxa more closely related to Heloderma than Varanus. It includes Heloderma, as well as several extinct genera, such as Estesia, Primaderma and Gobiderma, but this group was found to be polyphyletic in the most recent and complete squamate phylogenetic analysis by Reeder et al. (2015).

Uncredited · CC BY-SA 2.5 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- AnguimorphaPBDB
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Amniota
Time the record covers
At least 78 million years on record.
The record covers at least 110.1 Mya to 31.8 Mya. No occurrence read is dated outside 113.2 Mya to 15.98 Mya.
67 occurrences of Monstersauria on record, most of them in the Late Cretaceous. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
67 occurrences- Early Cretaceous1%
- Late Cretaceous85%
- Paleocene1%
- Eocene8%
- Oligocene4%
- Mioceneunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
20 formations
- Oldman Formation · Belly River Group15
- Lance Formation7
- Baruungoyot Formation6
- Hell Creek Formation · Montana Group5
- Cedar Mountain Formation · Dakota Group4
- Djadokhta Formation4
Loading coastlines…
The named rock units Monstersauria has been recovered from, largest first. Drawn where the rock is now, not where it formed. 6 occurrences have no formation on record, which is common for older literature.
Monstersauria and its relatives
Late CretaceousWhere Monstersauria and its relatives were recorded in the Late Cretaceous.
Found in the same rock
738 taxa- Champsosaurus25 sites
- Trionychidae25 sites
- Anura23 sites
- Scapherpeton tectum23 sites
- Baenidae22 sites
- Hadrosauridae22 sites
- Opisthotriton kayi22 sites
- Palaeosaniwa canadensis22 sites
- Ceratopsidae21 sites
- Lepisosteus occidentalis20 sites
- Chelydridae19 sites
- Crocodylia19 sites
- Myledaphus bipartitus18 sites
- Paronychodon18 sites
- Teleostei18 sites
- Tyrannosauridae17 sites
- Aves15 sites
- Estesesox foxi15 sites
- Holostei15 sites
- Multituberculata15 sites
- Odaxosaurus piger15 sites
- Coriops amnicolus14 sites
- Saurornitholestes langstoni14 sites
- Ankylosauridae13 sites
The Paleobiology Database records 59 sites for Monstersauria, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
30 references- Vertebrate microsite assemblages (exclusive of mammals) from the Foremost and Oldman Formations of the Judith River Group (Campanian) of southeastern Alberta: an illustrated guide15
- Evolution des lézards du Paléogène en Europe5
- Taxonomic composition and systematics of Late Cretaceous lizard assemblages from Ukhaa Tolgod and adjacent localities, Mongolian Gobi Desert5
- Fossil lizards of North America5
- A new taxon of helodermatid-like lizard from the Albian-Cenomanian of Utah4
- Fossil vertebrates from the Late Cretaceous Lance Formation, eastern Wyoming4
- Taxonomy and evolution of Late Cretaceous lizards (Reptilia: Squamata) from western Canada3
- Fossil Vertebrates from the Cerro del Pueblo Formation, Coahuila, Mexico, and the Distribution of Late Campanian (Cretaceous) Terrestrial Vertebrate Faunas2
- Braincase and phylogenetic relationships of Estesia mongoliensis from the Late Cretaceous of the Gobi Desert and the recognition of a new clade of lizards2
- Anguimorphans and related lizards from the Late Cretaceous of the Gobi Desert, Mongolia2
- The paleontology, geology and taphonomy of the Tooth Draw Deposit; Hell Creek Formation (Maastrictian), Butte County, South Dakota1
- The first lizard fossil (Reptilia: Squamata) from the Mesozoic of South Korea1
Each of the 67 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 18 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Monstersauria
- Source
- Wikipedia/Commons
- Credit
- nobody named on the record
- Licence
- CC BY-SA 2.5
- Commercial use
- the licence allows it
No 3D scan of Monstersauria was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itAnguimorpha through timeFormations of the Late CretaceousThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Norell and Gao 1997