Macraucheniidae
Macraucheniidae is a family in the extinct South American ungulate order Litopterna, that resembled camelids. They had three functional digits on the fore and hind feet, as well as elongate necks. The family is generally divided up into two subfamilies, Cramaucheniinae and Macraucheniinae. The family shows retraction of the nasal region, most extremely to the top of the skull in derived macraucheniine taxa like Macrauchenia. which has been interpreted to have supported a proboscis, perhaps like that of a saiga antelope to filter dust, or a moose-like prehensile lip. The earliest unambiguous members of the family date to the late Oligocene around 30 million years ago. Polymorphis from the Eocene has historically been placed as a macraucheniid, but this has been doubted. Most early representatives had a body masses in the range of 80–120 kilograms (180–260 lb), though some like Llullataruca were as small as 35–55 kilograms (77–121 lb), and the last representatives of the family from the Pleistocene like Macrauchenia were over 1,000 kilograms (2,200 lb). The family reached its apex of diversity during the late Miocene around 10-6 million years ago, before declining to only a few species belong to the genera Macrauchenia and Xenorhinotherium by the Late Pleistocene.

Ivan Iofrida · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- LitopternaPBDB
- Diet
- herbivorePBDB, for Litopterna
- Life habit
- scansorialPBDB, for Eutheria
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Eutheria
Time the record covers
At least 55 million years on record.
The record covers at least 56 Mya to 11.7 ka. No occurrence read is dated outside 57 Mya to 0 Mya.
231 occurrences of Macraucheniidae on record, most of them in the Miocene. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
231 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
69 formations
- Santa Cruz Formation13
- Sarmiento Formation11
- Cerro Azul Formation10
- Ituzaingo Formation7
- Monte Hermoso Formation7
- Epecuén Formation6
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The named rock units Macraucheniidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 45 further formations hold fewer occurrences each and are not listed. 77 occurrences have no formation on record, which is common for older literature.
Macraucheniidae and its relatives
MioceneWhere Macraucheniidae and its relatives were recorded in the Miocene.
Found in the same rock
1,662 taxa- Macrauchenia patagonica31 sites
- Macrauchenia19 sites
- Smilodon populator17 sites
- Equus16 sites
- Hippidion16 sites
- Notiomastodon platensis16 sites
- Promacrauchenia16 sites
- Glyptodon clavipes15 sites
- Megatherium americanum15 sites
- Theosodon15 sites
- Toxodon15 sites
- Equus caballus14 sites
- Panochthus14 sites
- Toxodon platensis14 sites
- Glyptodon reticulatus13 sites
- Panthera onca13 sites
- Gomphotheriidae12 sites
- Lestodon armatus12 sites
- Glossotherium11 sites
- Glyptodon11 sites
- Lama11 sites
- Pampatherium11 sites
- Panochthus tuberculatus11 sites
- Protypotherium11 sites
The Paleobiology Database records 161 sites for Macraucheniidae, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
130 references- Updated data on Litopterna from the Huayquerian Stage/Age (Late Miocene-Early Pliocene) of central-east Argentina11
- The Eocene to Pleistocene vertebrates of Bolivia and their stratigraphic context: a review11
- The beginning of the age of mammals in South America. Part I10
- Paleobiology of Santacrucian native ungulates (Meridiungulata: Astrapotheria, Litoptera and Notoungulata)6
- Miocene vertebrates from Entre Rios province, eastern Argentina6
- Advances in the understanding of Neogene mammalian fauna in the Pampean region (central Argentina) through revising “biozone” hypotheses based on new dates and biochronological analyses5
- Two new macraucheniids (Mammalia: Litopterna) from the late middle Miocene (Laventan South American Land Mammal Age) of Quebrada Honda, Bolivia5
- Biostratigraphy and biochronology of the Monte Hermoso Formation (early Pliocene) at its type locality, Buenos Aires Province, Argentina5
- Proterotheriidae and Macraucheniidae (Litopterna, Mammalia) from the Pleistocene of Rio Grande do Sul State, Brazil5
- Pleistocene mammals from Uruguay: Biostratigraphic, biogeographic and environmental connotations5
- La faune de vertébrés du Pléistocéne supérieur de l’aire archéologique de São Raimundo Nonato (Piauí, Brésil)5
- Late Cenozoic vertebrates from the Southern Pampean region: systematic and bio-chronostratigraphic update4
Each of the 231 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 118 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Macraucheniidae
- Source
- Wikipedia/Commons
- Credit
- Ivan Iofrida
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan was found. Smithsonian Open Access, MorphoSource and Sketchfab were all searched.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itLitopterna through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Ameghino 1889