Isoetes
Isoetes, commonly known as the quillworts, is a genus of lycopod. It is the only living genus in the family Isoetaceae and order Isoetales. As of 2016, there were about 200 recognized species, with a cosmopolitan distribution mostly in aquatic habitats but with the individual species often scarce to rare. Species virtually identical to modern quillworts have existed since the Jurassic epoch, though the timing of the origin of modern Isoetes is subject to considerable uncertainty.

Pattavina, Pete · Public Domain · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- quillwortPBDB
- Sits inside
- IsoetaceaePBDB
- Diet
- "photoautotroph"PBDB, for Streptophyta
- Environment
- freshwater,terrestrialPBDB, for Lycopodiophyta
Time the record covers
At least 121 million years on record.
The record covers at least 121.4 Mya to 11.7 ka. No occurrence read is dated outside 143.1 Mya to 0 Mya.
85 occurrences of Isoetes on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
85 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
4 formations
- Cullen Formation2
- Bhuj Formation2
- Renova Formation · Ruby River Basin Group1
- Weald Clay Formation · Wealden Group1
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The named rock units Isoetes has been recovered from, largest first. Drawn where the rock is now, not where it formed. 79 occurrences have no formation on record, which is common for older literature.
Isoetes and its relatives
PleistoceneWhere Isoetes and its relatives were recorded in the Pleistocene.
Found in the same rock
906 taxa- Poaceae69 sites
- Chenopodiaceae65 sites
- Ericaceae60 sites
- Quercus51 sites
- Pinus50 sites
- Taraxacum49 sites
- Aster45 sites
- Artemisia42 sites
- Cupressaceae42 sites
- Plantago42 sites
- Apiaceae41 sites
- Cyperaceae40 sites
- Brassicaceae37 sites
- Olea33 sites
- Ephedra29 sites
- Anthemis27 sites
- Asteraceae24 sites
- Caryophyllaceae24 sites
- Podocarpidites23 sites
- Botryococcus22 sites
- Cyathea22 sites
- Eucalyptus22 sites
- Monolete22 sites
- Casuarinaceae21 sites
The Paleobiology Database records 85 sites for Isoetes, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
19 references- Early Pleistocene vegetation change in upland south-eastern Australia21
- Site U138911
- Site U138610
- Site U13858
- Site U13878
- Site U13917
- Site U13906
- A diverse late-glacial (Mackinaw Phase) biota from Leamington, Ontario2
- A Catalogue of Fossil Plants from India Part 2: Palaeozoic and Mesozoic Megafossils2
- Site U13881
- The palaeoenvironment associated with a partial Iguanodon skeleton from the Upper Weald Clay (Barremian, Early Cretaceous) at Smokejacks Brickworks (Ockley, Surrey, UK), based on palynomorphs and ostracods1
- Miocene algae and spores from Tierra del Fuego, Argentina1
Each of the 85 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 7 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Isoetes
- Source
- Wikipedia/Commons
- Credit
- Pattavina, Pete
- Licence
- Public Domain
- Commercial use
- the licence allows it
No 3D scan of Isoetes was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Isoetes”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itIsoetaceae through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Linnaeus 1753