Casuarinaceae
The Casuarinaceae are a family of dicotyledonous flowering plants placed in the order Fagales, consisting of four genera and 91 species of trees and shrubs native to eastern Africa, Australia, Southeast Asia, Malesia, Papuasia, and the Pacific Islands. At one time, all species were placed in the genus Casuarina. Lawrence Alexander Sidney Johnson separated out many of those species and renamed them into the new genera of Gymnostoma in 1980 and 1982, Allocasuarina in 1982, and Ceuthostoma in 1988, with some additional formal descriptions of new species in each other genus. At the time, it was somewhat controversial. The monophyly of these genera was later supported in a 2003 phylogenetic study of the family. In the Wettstein system, this family was the only one placed in the order Verticillatae. Likewise, in the Engler, Cronquist, and Kubitzki systems, the Casuarinaceae were the only family placed in the order Casuarinales.

Marshman at English Wikipedia / Eric Guinther · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- FagalesPBDB
- Diet
- "photoautotroph"PBDB, for Angiospermae
Time the record covers
At least 72 million years on record.
The record covers at least 72.2 Mya to 11.7 ka. No occurrence read is dated outside 100.5 Mya to 0 Mya.
214 occurrences of Casuarinaceae on record, most of them in the Eocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
214 occurrences- Late Cretaceous8%
- Paleocene15%
- Eocene37%
- Oligocene3%
- Miocene12%
- Pleistocene23%
- Holoceneunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
24 formations
- Yaw Formation24
- Black Mingo Formation14
- Peedee Formation9
- Aquia Formation9
- Huitrera Formation8
- McNairy Formation8
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The named rock units Casuarinaceae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 97 occurrences have no formation on record, which is common for older literature.
Casuarinaceae and its relatives
EoceneWhere Casuarinaceae and its relatives were recorded in the Eocene.
Found in the same rock
1,075 taxa- Proteaceae52 sites
- Podocarpus49 sites
- Myrtaceae48 sites
- Casuarinidites46 sites
- Araucaria41 sites
- Eucalyptus41 sites
- Dacrydium37 sites
- Agathis36 sites
- Asteraceae36 sites
- Cunoniaceae36 sites
- Poaceae36 sites
- Carya35 sites
- Momipites microfoveolatus35 sites
- Acacia33 sites
- Chenopodiaceae33 sites
- Elaeocarpus33 sites
- Momipites33 sites
- Monolete33 sites
- Proteacidites33 sites
- Rapanea33 sites
- Cyperaceae32 sites
- Dodonaea32 sites
- Podocarpidites32 sites
- Cyathea30 sites
The Paleobiology Database records 186 sites for Casuarinaceae, and this reads all of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
36 references- Paleogene Sporomorph Biostratigraphy, Northeastern Virginia29
- Taxonomy and Biostratigraphy of Late Cretaceous and Paleogene Triatriate Pollen from South Carolina26
- A proto-monsoonal climate in the late Eocene of Southeast Asia: evidence from a sedimentary record in central Myanmar24
- Early Pleistocene vegetation change in upland south-eastern Australia22
- Ecology of Paleocene & Eocene vegetation at Kakahu, South Canterbury, New Zealand13
- A complete pollen record of the last 230 ka from Lynch's Crater, north-eastern Australia13
- Middle Miocene palaeotopography at Little Bay, near Maroubra, New South Wales12
- Biostratigraphy and paleoecology of lower Paleozoic, upper Cretaceous, and lower Tertiary rocks in U.S. Geological Survey New Madrid test wells, Southeastern Missouri9
- A Late Pleistocene and Holocene pollen diagram from Lynch’s Crater, northeastern Queensland, Australia9
- Casuarinaceae from the Eocene of Patagonia, Argentina8
- Palynology of the early Miocene Foulden Maar, Otago, New Zealand: Diversity following destruction5
- The Oligo-Miocene coal floras of southeastern Australia5
Each of the 214 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 24 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Casuarinaceae
- Source
- Wikipedia/Commons
- Credit
- Marshman at English Wikipedia / Eric Guinther
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
No 3D scan of Casuarinaceae was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itFagales through timeFormations of the EoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Brown 1814