Heterostraci
Heterostraci is an extinct subclass of pteraspidomorph, ostracoderm, jawless vertebrate that lived primarily in marine and estuary environments. Heterostraci existed from the mid-Ordovician to the conclusion of the Devonian.

Apokryltaros · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- PteraspidomorphaPBDB
- Motility
- actively mobilePBDB, for Chordata
Time the record covers
At least 109 million years on record.
The record covers at least 432.9 Mya to 323.4 Mya. No occurrence read is dated outside 443.1 Mya to 298.9 Mya.
784 occurrences of Heterostraci on record, most of them in the Devonian. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
784 occurrences- Silurian15%
- Devonian84%
- Carboniferousunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
118 formations
- Delorme Formation57
- Peel Sound Formation43
- Dniester Formation39
- Ben Nevis Formation37
- Kureika Formation36
- Fraenkelryggen Formation31
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The named rock units Heterostraci has been recovered from, largest first. Drawn where the rock is now, not where it formed. 94 further formations hold fewer occurrences each and are not listed. 98 occurrences have no formation on record, which is common for older literature.
Heterostraci and its relatives
DevonianWhere Heterostraci and its relatives were recorded in the Devonian.
Found in the same rock
1,193 taxa- Cephalaspis20 sites
- Corvaspis20 sites
- Protopteraspis gosseleti19 sites
- Traquairaspis18 sites
- Acanthodii15 sites
- Glyptolepis14 sites
- Cyathaspididae13 sites
- Poraspis13 sites
- Turinia pagei13 sites
- Nikolivia elongata12 sites
- Cheirolepis11 sites
- Byssacanthus dilatatus10 sites
- Protopteraspis10 sites
- Actinolepis tuberculata9 sites
- Bivalvia9 sites
- Ostracoda9 sites
- Poraspis polaris9 sites
- Asterolepis estonica8 sites
- Kujdanowiaspis podolica8 sites
- Nostolepis8 sites
- Orvikuina8 sites
- Phialaspis symondsi8 sites
- Podolaspis lerichei8 sites
- Psammosteus8 sites
The Paleobiology Database records 286 sites for Heterostraci, and this reads 240 of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
123 references- The Early Devonian armoured agnathans of Podolia, Ukraine71
- Upper Silurian and Devonian heterostracan pteraspidomorphs (Vertebrata) from Severnaya Zemlya (Russia): a preliminary report with biogeographical and biostratigraphical implications63
- The vertebrate stratigraphy of the Lower Devonian (Red Bay Group and Wood Bay Formation) of Spitsbergen58
- Ostracoderm faunas of the Delorne and associated Siluri-Devonian Formations, North Western Territories, Canada56
- The Middle Devonian fishes of the Baltic States (Estonia, Latvia) and Belarus47
- Subdivision of the Kureyka suite based on amphiaspid (Agnatha) assemblages43
- Habitat of Llandoverian-Lochkovian eurypterids29
- A New Heterostracan from the Upper Silurian of Northwest Territories, Canada26
- A new subfamily of the Pteraspididae (Agnatha, Heterostraci) from the Upper Silurian and Lower Devonian of Arctic Canada25
- Protopteraspis Gosseleti (Vertebrata: Pteraspidomorphi: Heterostraci) from the Lower Devonian of Shropshire, England21
- Review of the Pteraspidiform heterostracans (Vertebrata, Agnatha) from the Devonian of Podolia, Ukraine, in the Theodor Vascautanu Collection, Bucharest, Romania18
- A new species of the Devonian actinopterygian fish Moythomasia from Belarus14
Each of the 784 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 111 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Heterostraci
- Source
- Wikipedia/Commons
- Credit
- Apokryltaros
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan of Heterostraci was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itPteraspidomorpha through timeFormations of the DevonianThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Lankester 1868)