Cydnidae
Cydnidae are a family of pentatomoid bugs, known by common names including burrowing bugs or burrower bugs. As the common name would suggest, many members of the group live a subterranean lifestyle, burrowing into soil using their head and forelegs, only emerging to mate and then laying their eggs in soil. Other members of the group are not burrowers, and live above the soil layer, often in close association with plants. Several species are known as agricultural pests.

Didier Descouens · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- burrowing bugPBDB
- Sits inside
- PentatomoideaPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 123 million years on record.
The record covers at least 125.77 Mya to 2.58 Mya. No occurrence read is dated outside 132.6 Mya to 1.8 Mya.
145 occurrences of Cydnidae on record, most of them in the Oligocene. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
145 occurrences- Early Cretaceous8%
- Late Cretaceous6%
- Eocene30%
- Oligocene37%
- Miocene19%
- Plioceneunder 1%
- Pleistoceneunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
24 formations
- Florissant Formation20
- Niveau du gypse d'Aix Formation11
- Salt Formation10
- Upper Freshwater-Molasse Formation8
- Eifel Formation8
- Green River Formation5
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The named rock units Cydnidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 50 occurrences have no formation on record, which is common for older literature.
Cydnidae and its relatives
OligoceneWhere Cydnidae and its relatives were recorded in the Oligocene.
Found in the same rock
3,680 taxa- Curculionidae13 sites
- Coleoptera12 sites
- Apioninae10 sites
- Cydnus brevicollis10 sites
- Formicidae10 sites
- Plecia10 sites
- Lygaeidae8 sites
- Pentatomidae8 sites
- Plecia bucklandi8 sites
- Plecia foersteri8 sites
- Acrididae7 sites
- Aphanus pulchellus7 sites
- Protainophthalmus asperulus7 sites
- Entimini6 sites
- Libellulidae6 sites
- Plecia livida6 sites
- Staphylinidae6 sites
- Bibionidae5 sites
- Cydnopsis haidingeri5 sites
- Cydnus armiger5 sites
- Cydnus tertiarius5 sites
- Diptera5 sites
- Ichneumonidae5 sites
- Mycetophilidae5 sites
The Paleobiology Database records 66 sites for Cydnidae, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
39 references- Die Insekten der "Plattigen Steinmergels" von Brunstatt23
- The Tertiary Insects of North America22
- Les insectes fossiles des terrains oligocènes de France17
- Heer, 185315
- Systematik, Phylogenie, Taphonomie und Paläoökologie der Insekten aus dem Mittel-Eozän des Eckfelder Maares, Vulkaneifel8
- Peloridiina (= Coleorrhyncha) and Cimicina (= Heteroptera), in Nasekomye v rannemelovykh ekosistemakh zapadnoy Mongolii5
- Geocorisae (Landwanzen) aus den Oberoligocanen Ablagerungen von Rott5
- Morphological phylogeny of new Cretaceous fossils elucidates the early history of soil dwelling among bugs4
- Fossil Cydnidae (Heteroptera) in the Dominican Amber4
- Nouvelles recherches sur quelques insectes des Plâtrières d'Aix en Provence3
- The Fossil Insects of the Green River Shales3
- Sur un Hemiptère de la famille des Pentatomides3
Each of the 145 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 27 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Cydnidae
- Source
- Wikipedia/Commons
- Credit
- Didier Descouens
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
No 3D scan of Cydnidae was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Cydnidae”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itPentatomoidea through timeFormations of the OligoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Billberg 1820)