Colobinae
The Colobinae or leaf-eating monkeys are a subfamily of the Old World monkey family that includes 61 species in 11 genera, including the black-and-white colobus, the large-nosed proboscis monkey, and the gray langurs. Some classifications split the colobine monkeys into two tribes, while others split them into three groups. Both classifications put the three African genera Colobus, Piliocolobus, and Procolobus in one group; these genera are distinct in that they have stub thumbs, hence their family name, which comes from Ancient Greek κολοβός (kolobós), meaning "docked". The various Asian genera are placed into another one or two groups. Analysis of mtDNA confirms the Asian species form two distinct groups, one of langurs and the other of the "odd-nosed" species, but are inconsistent as to the relationships of the gray langurs; some studies suggest that the gray langurs are not closely related to either of these groups, while others place them firmly within the langur group.

Dr. Ajay B. · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- CercopithecidaePBDB
- Diet
- frugivore, folivorePBDB, for Cercopithecidae
- Life habit
- arborealPBDB, for Cercopithecidae
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Eutheria
Time the record covers
At least 11 million years on record.
The record covers at least 11.63 Mya to 11.7 ka. No occurrence read is dated outside 23.04 Mya to 0 Mya.
254 occurrences of Colobinae on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
254 occurrences- Miocene12%
- Pliocene32%
- Pleistocene55%
- Holoceneunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
37 formations
- Shungura Formation · Omo Group80
- Koobi Fora Formation · Omo Group28
- Vogel River Series Formation14
- Nachukui Formation · Omo Group6
- USNO Formation · Omo Group5
- Adu-Asa Formation · Awash Group4
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The named rock units Colobinae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 13 further formations hold fewer occurrences each and are not listed. 64 occurrences have no formation on record, which is common for older literature.
Colobinae and its relatives
PleistoceneWhere Colobinae and its relatives were recorded in the Pleistocene.
Found in the same rock
1,388 taxa- Hipparion67 sites
- Gazella50 sites
- Kolpochoerus limnetes48 sites
- Rhinocolobus turkanaensis47 sites
- Palaeoloxodon recki46 sites
- Hominidae44 sites
- Equus38 sites
- Alcelaphini37 sites
- Bovidae37 sites
- Ceratotherium simum36 sites
- Paracolobus mutiwa35 sites
- Cercopithecidae32 sites
- Hystrix32 sites
- Gerontochoerus scotti30 sites
- Tragelaphus nakuae30 sites
- Hylochoerus euilus29 sites
- Kolpochoerus27 sites
- Libytherium maurusium27 sites
- Adenota recki26 sites
- Metridiochoerus25 sites
- Kobus sigmoidalis24 sites
- Reduncini24 sites
- Theropithecus oswaldi24 sites
- Tragelaphus24 sites
The Paleobiology Database records 193 sites for Colobinae, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
100 references- Extinct large Colobines from the Plio-Pleistocene of Africa61
- The Omo Group: Archives of the International Omo Research Expedition20
- Laetoli: a Pliocene Site in Northern Tanzania14
- Lothagam: the dawn of humanity in eastern Africa10
- Pleistocene fossil Primates from Luoding, Guangdong9
- The Omo Micromammals: Systematics and Paleoecology of Early Man Sites from Ethiopia6
- Mandibular postcanine dentition from the Shungura Formation, Ethiopia: Crown morphology, taxonomic allocations and Plio-Pleistocene Hominid Evolution5
- Late Miocene mammal localities of the lower Axios valley (Macedonia, Greece) and their stratigraphic significance4
- Quaternary langurs and macaques from the Malay Archipelago4
- Large Ruscinian Mammalia in the Territory of the Former Soviet Union3
- Refined stratigraphy of the hominid-bearing Awash Group, Middle Awash Valley, Afar Depression, Ethiopia3
- Pliocene and Pleistocene mammalian biostratigraphy in southeastern Transylvania (Romania)3
Each of the 254 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 88 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Colobinae
- Source
- Wikipedia/Commons
- Credit
- Dr. Ajay B.
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
No 3D scan was found. Smithsonian Open Access, MorphoSource and Sketchfab were all searched.
The description on this site
- Source
- Wikipedia, “Colobinae”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itCercopithecidae through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Jerdon 1867