Bethylidae
The Bethylidae are a family of aculeate wasps in the superfamily Chrysidoidea. As a family, their biology ranges between parasitoid wasps and hunting wasps.

This image is created by user B. Schoenmakers at Waarneming.nl , a source of nature observations in the Netherlands. · CC BY 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- flat waspPBDB
- Sits inside
- ChrysidoideaPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 121 million years on record.
The record covers at least 121.4 Mya to 11.7 ka. No occurrence read is dated outside 125.77 Mya to 0 Mya.
144 occurrences of Bethylidae on record, most of them in the Eocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
144 occurrences- Early Cretaceous8%
- Late Cretaceous20%
- Eocene60%
- Oligoceneunder 1%
- Miocene8%
- Holocene3%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
12 formations
- Argiles d'lignite du Soissonnais Formation11
- Kheta Formation5
- Escucha Formation4
- Florissant Formation2
- Foremost Formation · Belly River Group2
- la Pedrera de Rubies Formation1
Loading coastlines…
The named rock units Bethylidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 113 occurrences have no formation on record, which is common for older literature.
Bethylidae and its relatives
EoceneWhere Bethylidae and its relatives were recorded in the Eocene.
Found in the same rock
4,107 taxa- Prenolepis henschei7 sites
- Yantaromyrmex geinitzi7 sites
- Gesomyrmex hoernesi6 sites
- Nylanderia pygmaea6 sites
- Tetraponera simplex6 sites
- Ctenobethylus goepperti5 sites
- Orchestina baltica5 sites
- Plagiolepis kuenowi5 sites
- Pseudoteutana stigmatosa5 sites
- Segestria5 sites
- Brachypogon (Isohelea) prominulus4 sites
- Ceratopogon4 sites
- Ceratopogon forcipiformis4 sites
- Coleoptera4 sites
- Culicoides4 sites
- Culicoides speciosus4 sites
- Dictynidae4 sites
- Eocryphoeca gracilipes4 sites
- Eohelea sinuosa4 sites
- Formica flori4 sites
- Formica gustawi4 sites
- Formicidae4 sites
- Gnamptogenys europaea4 sites
- Lasius schiefferdeckeri4 sites
The Paleobiology Database records 58 sites for Bethylidae, and this reads all of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
59 references- The Parasitic Hymenoptera of the Baltic Amber18
- Synopsis of the fossil flat wasps Epyrinae (Hymenoptera, Bethylidae), with description of three new genera and 10 new species11
- Revision of Bethylinae fossils (Hymenoptera: Bethylidae) from Baltic, Rovno and Oise amber, with comments on the Tertiary fauna of the subfamily11
- Zur Phylogenie und Systematik der Bethylidae (Insecta: Hymenoptera, Chrysidoidea)11
- Diversity and phylogeny of the extinct wasp subfamily Lancepyrinae (Hymenoptera, Bethylidae) revealed by mid-Cretaceous Burmese amber6
- Synopsis of the fossil Pristocerinae (Hymenoptera, Bethylidae), with description of two new genera and six species from Burmese, Taimyr, Baltic and Rovno ambers6
- Some New Fossil Parasitic Hymenoptera from Baltic Amber6
- Synopsis of the fossil Scleroderminae (Hymenoptera, Bethylidae) with description of a new genus and four new species from Baltic amber4
- Bethylidae from Early Cretaceous Spanish amber (Hymenoptera: Chrysidoidea)4
- Cretaceous aculeate wasps from Taimyr, Siberia (Hymenoptera)4
- The first flat wasps (Hymenoptera: Bethylidae) from Miocene Mexican amber3
- Eocene bethylid wasps from French amber (Hymenoptera: Bethylidae)3
Each of the 144 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 47 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Bethylidae
- Source
- Wikipedia/Commons
- Credit
- This image is created by user B. Schoenmakers at Waarneming.nl , a source of nature observations in the Netherlands.
- Licence
- CC BY 3.0
- Commercial use
- the licence allows it
No 3D scan of Bethylidae was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Bethylidae”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itChrysidoidea through timeFormations of the EoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Forster 1856