Culicoides
Culicoides is a genus of biting midges in the family Ceratopogonidae. There are over 1000 species in the genus, which is divided into many subgenera. Several species are known to be vectors of various diseases and parasites which can affect animals. The genus has a long fossil record, with earliest known fossils being from Burmese amber, around 99 million years old.

Scott Bauer, USDA ARS · Public Domain · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- CulicoidiniPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 75 million years on record.
The record covers at least 93.9 Mya to 18.5 Mya. No occurrence read is dated outside 100.5 Mya to 13.82 Mya.
116 occurrences of Culicoides on record, most of them in the Late Cretaceous. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
116 occurrences- Late Cretaceous44%
- Eocene33%
- Oligocene10%
- Miocene13%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
6 formations
- Foremost Formation · Belly River Group20
- Rott Formation12
- Raritan Formation · Potomac Group7
- Barstow Formation5
- Kheta Formation5
- Mazantic Shale Formation · Simojovel Group1
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The named rock units Culicoides has been recovered from, largest first. Drawn where the rock is now, not where it formed. 66 occurrences have no formation on record, which is common for older literature.
Culicoides and its relatives
Late CretaceousWhere Culicoides and its relatives were recorded in the Late Cretaceous.
Found in the same rock
3,743 taxa- Ceratopogon12 sites
- Forcipomyia12 sites
- Eohelea sinuosa11 sites
- Forcipomyia (Forcipomyia)11 sites
- Monogedania clunipes11 sites
- Atriculicoides globosus10 sites
- Brachypogon (Isohelea) prominulus9 sites
- Serromyia9 sites
- Brachypogon (Isohelea)8 sites
- Ceratopogon forcipiformis8 sites
- Dasyhelea8 sites
- Prenolepis henschei8 sites
- Serphites paradoxus8 sites
- Tetraponera simplex8 sites
- Yantaromyrmex geinitzi8 sites
- Ctenobethylus goepperti7 sites
- Gesomyrmex hoernesi7 sites
- Lasius schiefferdeckeri7 sites
- Meunierohelea7 sites
- Nannohelea7 sites
- Nylanderia pygmaea7 sites
- Pachycondyla succinea7 sites
- Peronehelea chrimikalydia7 sites
- Plagiolepis klinsmanni7 sites
The Paleobiology Database records 44 sites for Culicoides, and this reads all of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
22 references- Borkent, 199527
- Biting midges (Diptera, Ceratopogonidae) from Baltic amber25
- Neue Dipteren (Nematocera) aus dem Oberoligocan von Rott. 1. Teil 6. Familie: Tendipedidae (Zuck- oder Schwarmmucken). 2. Teil 7. Familie: Heleidae (Gnitzen) 3. Teil 8. Familie: Lycoriidae (Trauermucken)12
- Biting midges from Dominican amber. III. Species of the tribes Culicoidini and Ceratopogonini (Diptera: Ceratopogonidae)9
- Biting midges (Diptera, Ceratopogonidae) from Miocene Saxonian amber6
- Biting midges (Diptera: Ceratopogonidae) in Eocene Baltic amber from the Rovno region (Ukraine)5
- Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous4
- Further biting midges (Diptera: Ceratopogonidae) in Canadian Cretaceous amber4
- Late Cretaceous biting midges (Diptera, Ceratopogonidae) from fossil resins of the Khatanga Depression4
- Further biting midges (Diptera: Ceratopogonidae) from Upper Cretaceous New Jersey amber3
- Biting Midges from Upper Cretaceous New Jersey Amber (Diptera: Ceratopogonidae)3
- Miocene arthropods from the Mojave desert, California3
Each of the 116 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 10 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Culicoides
- Source
- Wikipedia/Commons
- Credit
- Scott Bauer, USDA ARS
- Licence
- Public Domain
- Commercial use
- the licence allows it
No 3D scan was found. Smithsonian Open Access, MorphoSource and Sketchfab were all searched.
The description on this site
- Source
- Wikipedia, “Culicoides”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itCulicoidini through timeFormations of the Late CretaceousThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Latreille 1809