Artiodactyla
Artiodactyla is an order of placental mammals. Typically, they are ungulates which bear weight equally on two of their five toes. The other three toes are either present, absent, vestigial, or pointing posteriorly. By contrast, most perissodactyls bear weight on an odd number of the five toes. Another difference between the two orders is that many artiodactyls digest plant cellulose in one or more stomach chambers rather than in their intestine. Molecular biology, along with fossil discoveries, has found that cetaceans fall within this taxonomic branch, being most closely related to hippopotamuses. Some modern taxonomists thus apply the name Cetartiodactyla to this group, while others opt to include cetaceans within the existing name of Artiodactyla. Some researchers use "even-toed ungulates" to exclude cetaceans and only include terrestrial artiodactyls, making the term paraphyletic in nature.

LittleJerry · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- even-toed ungulatePBDB
- Sits inside
- ArtiodactylamorphaPBDB
- Diet
- grazer, browserPBDB
- Life habit
- ground dwellingPBDB
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB
Time the record covers
At least 51 million years on record.
The record covers at least 51.1 Mya to 11.7 ka. No occurrence read is dated outside 56 Mya to 0 Mya.
12,000+ occurrences of Artiodactyla on record. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
12,000 occurrences readThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. The read hit its cap, so every figure above is a floor and the shares are of the occurrences read. More on what this infers.
Found in
at least 387 formations
- Koobi Fora Formation912
- Shungura Formation · Omo Group661
- Brule Formation · White River Group494
- Nachukui Formation · Omo Group422
- Vogel River Series Formation234
- Valentine Formation · Ogallala Group198
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The named rock units Artiodactyla has been recovered from, largest first. Drawn where the rock is now, not where it formed. The read hit its cap, so each count is a floor. 363 further formations hold fewer occurrences each and are not listed. 3,410 occurrences have no formation on record, which is common for older literature.
Found in the same rock
1,259 taxa- Hyopsodus99 sites
- Coryphodon80 sites
- Diacodexis73 sites
- Phenacodus vortmani60 sites
- Paramys excavatus56 sites
- Paramys copei53 sites
- Haplomylus speirianus50 sites
- Phenacolemurinae50 sites
- Sifrhippus grangeri49 sites
- Phenacodus48 sites
- Didymictis47 sites
- Esthonyx bisulcatus45 sites
- Prototomus41 sites
- Hyracotherium38 sites
- Cantius34 sites
- Prodiacodon tauricinerei34 sites
- Vulpavus australis34 sites
- Ectocion osbornianus32 sites
- Viverravus32 sites
- Bunophorus31 sites
- Microsyopidae31 sites
- Cantius abditus30 sites
- Apheliscus29 sites
- Esthonyx spatularius29 sites
The Paleobiology Database records at least 866 sites for Artiodactyla, and this reads 240 of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
at least 1,061 references- Synonymies and reidentifications of North American fossil mammals1,080
- Family Bovidae544
- Stratigraphy and Paleontology of pliocene and Pleistocene Localities West of Lake Turkana, Kenya395
- Les artiodactyles de l'Eocene moyen et superieur d'Europe occidentale; systematique et evolution255
- Laetoli: a Pliocene Site in Northern Tanzania234
- The Phylogeny of the Oreodonts. Part 1: Merycoidodontinae, Eporeodontinae, and Leptaucheniinae, Three Subfamilies Of Oreodonts, With an Appendix to the Revision Of The Merycoidodontidae, and Part 2. Summary and Conclusions Concerning the Merycoidodontidae181
- Family Suidae174
- The Omo Group: Archives of the International Omo Research Expedition164
- Extinct large Colobines from the Plio-Pleistocene of Africa154
- Vertebrate biostratigraphy of the Ogallala Group in Nebraska143
- The Omo Micromammals: Systematics and Paleoecology of Early Man Sites from Ethiopia126
- Paleoenvironmental and Cultural Implications of Late Holocene Archaeological Faunas from the Orange Free State and North Central Cape Province, South Africa120
Each of the 12,000 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. That read hit its cap, so each count is a floor. 1,049 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content3
Picture of Artiodactyla
- Source
- Wikipedia/Commons
- Credit
- LittleJerry
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
Photogrammetry
- Source
- MorphoSource
- Licence
- In Copyright
- Commercial use
- cannot be read from the licence given — check the record
- Specimen
- Artiodactyla indet. · msn_fir:mgp:IGF_105128
3D model
- Source
- Sketchfab
- Credit
- SchmalhausenEvolMorph
- Licence
- CC Attribution
- Commercial use
- cannot be read from the licence given — check the record
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itArtiodactylamorpha through timeThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Owen 1848)