Amara
Amara is a large genus of carabid beetles, commonly called the sun beetles. Many are holarctic, but a few species are neotropical or occur in eastern Asia.

Uncredited · Attribution · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- AmarinaPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 47 million years on record.
The record covers at least 48.07 Mya to 129 ka. No occurrence read is dated outside 56 Mya to 11.7 ka.
107 occurrences of Amara on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
107 occurrences- Eocene15%
- Oligocene2%
- Miocene6%
- Pleistocene78%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
11 formations
- Florissant Formation14
- Keremesit Formation14
- Kap København Formation11
- Edoma Formation7
- Oler Formation4
- Upper Freshwater-Molasse Formation3
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The named rock units Amara has been recovered from, largest first. Drawn where the rock is now, not where it formed. 47 occurrences have no formation on record, which is common for older literature.
Amara and its relatives
PleistoceneWhere Amara and its relatives were recorded in the Pleistocene.
Found in the same rock
1,704 taxa- Curtonotus alpinus41 sites
- Morychus viridis41 sites
- Pterostichus (Cryobius)40 sites
- Phyllobius37 sites
- Stephanocleonus eruditus33 sites
- Aphodius32 sites
- Poecilus nearcticus31 sites
- Bembidion25 sites
- Hypera ornata23 sites
- Chrysolina rufilabris22 sites
- Cholevinus sibiricus20 sites
- Stephanocleonus foveifrons19 sites
- Coniocleonus ferrugineus17 sites
- Troglocollops arcticus17 sites
- Lepyrus nordenskioldi14 sites
- Stenus14 sites
- Agabus13 sites
- Chrysolina subsulcata13 sites
- Chrysolina tolli13 sites
- Harpalus amputatus amputatoides13 sites
- Hydroporus13 sites
- Stephanocleonus fossulatus13 sites
- Tachinus arcticus12 sites
- Harpalus vittatus kiselevi11 sites
The Paleobiology Database records 70 sites for Amara, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
19 references- Late Cenozoic insects of northern Eurasia57
- Palaeoentomology of the Kap København Formation, a Plio-Pleistocene sequence in Peary Land, North Greenland11
- Adephagous and clavicorn Coleoptera from the Tertiary deposits at Florissant, Colorado with descriptions of a few other forms and a systematic list of the non-rhynchophorus Tertiary Coleoptera of North America8
- A Late Pleistocene fauna and flora from Upton Warren, Worcestershire7
- Marine Isotope Stage 9 environments of fluvial deposits at Hackney, north London, UK3
- Beiträge zur Insektenfauna Oeningens: Coleoptera. Geodephagen, Hydrocanthariden, Gyriniden, Brachelytren, Clavicornen, Lamellicornen und Buprestiden3
- Paleogeography of the lower Volga at the end of Atelian time based on entomological and botanical data2
- Fossil Coleoptera from Florissant in the United States National Museum2
- A report on some recent collections of fossil Coleoptera from the Miocene shales of Florissant2
- Pleistoceńskie owady z Borysławia2
- Die Insekten der "Plattigen Steinmergels" von Brunstatt2
- The paleoenvironment of the Olympia beds based on fossil beetles from Discovery Park, Seattle, Washington, U.S.A.1
Each of the 107 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 7 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content2
Picture of Amara
- Source
- Wikipedia/Commons
- Credit
- nobody named on the record
- Licence
- Attribution
- Commercial use
- cannot be read from the licence given — check the record
Photogrammetry
- Source
- MorphoSource
- Licence
- not stated by the source
- Commercial use
- cannot be read from the licence given — check the record
- Specimen
- Amara Amara plebeja · ETHZ:ENT:0576136
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itAmarina through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Bonelli 1810