Tornoceratidae
Tornoceratidae is a family of goniatitid ammonoids from the middle and upper Devonian. The family is included in the suborder Tornoceratina and the superfamily Tornoceratoidea.

Not Tornoceratidae. This is Tornoceras uniangulare (Conrad, 1842), a relative — both are Tornoceratidae.
Yale Peabody Museum · CC0 · via GBIF. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- TornocerataceaePBDB
- Diet
- carnivorePBDB, for Ammonoidea
- Life habit
- nektonicPBDB, for Goniatitida
- Motility
- fast-movingPBDB, for Ammonoidea
- Vision
- well-developedPBDB, for Cephalopoda
- Environment
- marinePBDB, for Cephalopoda
Time the record covers
At least 22 million years on record.
The record covers at least 387.95 Mya to 365.2 Mya. No occurrence read is dated outside 393.47 Mya to 358.86 Mya.
442 occurrences of Tornoceratidae on record, most of them in the Devonian. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
442 occurrences- Devonian100%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
44 formations
- Virgin Hills Formation71
- Sand Formation53
- Domanik Formation39
- Lyaiol Formation34
- Cashaqua Formation · Sonyea Group17
- Angola Formation · West Falls Group14
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The named rock units Tornoceratidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 20 further formations hold fewer occurrences each and are not listed. 73 occurrences have no formation on record, which is common for older literature.
Tornoceratidae and its relatives
DevonianWhere Tornoceratidae and its relatives were recorded in the Devonian.
Found in the same rock
633 taxa- Tornoceras55 sites
- Tornoceras simplex44 sites
- Manticoceras lamed24 sites
- Tornoceras typum24 sites
- Polonoceras delepinei23 sites
- Sporadoceras23 sites
- Sporadoceras muensteri19 sites
- Eldredgeops rana15 sites
- Greenops boothi14 sites
- Manticoceras14 sites
- Manticoceras intumescens14 sites
- Manticoceras sinuosum14 sites
- Paratornoceras lentiforme14 sites
- Probeloceras lutheri14 sites
- Tornoceras uniangulare14 sites
- Mucrospirifer mucronatus13 sites
- Lobobactrites12 sites
- Manticoceras cordatum12 sites
- Chonetoidea11 sites
- Mediospirifer audaculus11 sites
- Acanthoclymenia neapolitana10 sites
- Ambocoelia10 sites
- Aulatornoceras10 sites
- Aulatornoceras eifliense10 sites
The Paleobiology Database records 289 sites for Tornoceratidae, and this reads 240 of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
50 references- Revision of ammonoid biostratigraphy in the Frasnian (Upper Devonian) of the Southern Timan (northeast Russian Platform)77
- Late Devonian goniatites (Cephalopoda, Ammonoidea) from New York State72
- Upper Devonian (Famennian) ammonoids from the Canning Basin, Western Australia58
- Upper Frasnian Tornoceratidae (Ammonoidea) from the Sand Formation (Bergisch Gladbach‑Pafrath Syncline, Rhenish Massif)53
- Devonskie ammonoidei I. Agoniatitidy32
- Devonian goniatite biostratigraphy and timing of facies movements in the Frasnian of the Canning Basin, Western Australia14
- Devonian (Frasnian) goniatites from Waterside Cove and Staverton Wood, south Devon10
- Definition of the Frasnian/Famennian Stage boundary10
- Redefinition, Stratigraphy and Depostional Environments of the Mottville Member (Hamilton Group) in Central and Eastern New York10
- Upper Devonian goniatites and co-occurring conodonts from the Holy Cross Mountains: studies of the Polish Geological Institute collections9
- Die Fauna aus dem mittleren Oberdevon (Nehden-Stufe) des suedwestlichen Bergischen Landes (Rheinisches Schiefergebirge); The lower Famennian fauna (Nehden-substage) of the southwestern region of the Bergisches Land (Rhenish Schiefergebirge)9
- The “Nehden Goniatite Shale” (lower Famennian, Brilon Reef Complex, NE Rhenish Massif)8
Each of the 442 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 38 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1

Picture of Tornoceras uniangulare (Conrad, 1842)
- Source
- GBIF
- Credit
- Yale Peabody Museum
- Licence
- CC0
- Commercial use
- the licence allows it
No 3D scan was found. Smithsonian Open Access, MorphoSource and Sketchfab were all searched.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itTornocerataceae through timeFormations of the DevonianThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Arthaber 1911