Saniwa
Saniwa is an extinct genus of varanid lizard that lived during the Eocene epoch. It is known from well-preserved fossils found in the Bridger and Green River Formations of Wyoming, United States. The type species S. ensidens was described in 1870 as the first fossil lizard known from North America. A second species, S. orsmaelensis, is recognised from remains found in Europe. It is a close relative of Varanus, the genus that includes monitor lizards.

Jonathan Chen · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- SaniwinaePBDB
- Diet
- carnivorePBDB
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB
Time the record covers
At least 27 million years on record.
The record covers at least 60.9 Mya to 33.9 Mya. No occurrence read is dated outside 63.4 Mya to 15.98 Mya.
71 occurrences of Saniwa on record, most of them in the Eocene. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
71 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
21 formations
- Bridger Formation18
- San Jose Formation8
- Uinta Formation5
- Friars Formation · La Jolla Group4
- Willwood Formation4
- Wind River Formation4
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The named rock units Saniwa has been recovered from, largest first. Drawn where the rock is now, not where it formed. 7 occurrences have no formation on record, which is common for older literature.
Saniwa and its relatives
EoceneWhere Saniwa and its relatives were recorded in the Eocene.
Found in the same rock
1,041 taxa- Glyptosaurinae14 sites
- Trionychidae11 sites
- Hyopsodus9 sites
- Iguanidae9 sites
- Anguidae8 sites
- Apatemys8 sites
- Boavus occidentalis7 sites
- Herpetotherium knighti7 sites
- Echmatemys6 sites
- Glyptosaurus sylvestris6 sites
- Hyrachyus modestus6 sites
- Lacertilia6 sites
- Parasauromalus olseni6 sites
- Viverravus minutus6 sites
- Antiacodon pygmaeus5 sites
- Anura5 sites
- Apternodontidae5 sites
- Boidae5 sites
- Gekkonidae5 sites
- Gerrhonotinae5 sites
- Helodermatidae5 sites
- Necrosaurus5 sites
- Omomys carteri5 sites
- Palaeoxantusia5 sites
The Paleobiology Database records 63 sites for Saniwa, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
29 references- Fossil lizards of North America13
- Fossil Squamata from the San José Formation, Early Eocene, San Juan Basin, New Mexico8
- Evolution des lézards du Paléogène en Europe7
- Summary of known occurrences of terrestrial vertebrates from Eocene strata of southern California5
- Preliminary description of new Tertiary reptiles. Part I and Part II5
- Amphibians and Reptiles (in: The Geology and Paleontology of the Elk Mountain and Tabernacle Butte area, Wyoming)4
- Preliminary analysis of the vertebrate fossil fauna of the Boysen Reservoir area3
- Early Eocene lizards of the Wasatch Formation near Bitter Creek, Wyoming: diversity and paleoenvironment during an interval of global warming2
- A new lizard assemblage from the earliest Eocene (zone Wa0) of the Bighorn Basin, Wyoming, USA: biogeography during the warmest interval of the Cenozoic2
- The anatomy of the fossil varanid lizard Saniwa ensidens Leidy, 1870, based on a newly discovered complete skeleton2
- Basin margins, biodiversity, evolutionary innovation, and the origin of new taxa2
- A diverse new assemblage of late Eocene squamates (Reptilia) from the Chadron Formation of North Dakota, U.S.A.1
Each of the 71 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 17 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content2
Picture of Saniwa
- Source
- Wikipedia/Commons
- Credit
- Jonathan Chen
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
CT scan
- Source
- MorphoSource
- Licence
- In Copyright
- Commercial use
- cannot be read from the licence given — check the record
- Specimen
- Saniwa sp. · amnh:farb:8688
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Saniwa”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itSaniwinae through timeFormations of the EoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Leidy 1870