Protopteraspididae
Protopteraspididae is an extinct family of pteraspidid heterostracan agnathans. Fossils of the various genera are found in early Devonian-aged marine strata. Protopteraspidids were once thought to represent a taxon of basal pteraspidids but recent evaluations demonstrate that Protopteraspididae is a paraphyletic group of various transitional forms representing a gradual transition between the more advanced Pteraspoidei, and the anchipteraspidids and the Psammosteids.

Apokryltaros · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- PteraspidiformesPBDB
- Motility
- actively mobilePBDB, for Chordata
Time the record covers
At least 23 million years on record.
The record covers at least 416.8 Mya to 393.47 Mya. No occurrence read is dated outside 419.62 Mya to 387.95 Mya.
65 occurrences of Protopteraspididae on record, most of them in the Devonian. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
65 occurrences- Devonian100%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
20 formations
- Psammosteus Limestone Formation18
- Ben Nevis Formation8
- Fraenkelryggen Formation5
- Wood Bay Formation5
- Tyver Formation5
- Spokojnaya Formation4
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The named rock units Protopteraspididae has been recovered from, largest first. Drawn where the rock is now, not where it formed.
Protopteraspididae and its relatives
DevonianWhere Protopteraspididae and its relatives were recorded in the Devonian.
Found in the same rock
273 taxa- Protopteraspis gosseleti19 sites
- Cephalaspis12 sites
- Poraspis10 sites
- Protopteraspis10 sites
- Corvaspis9 sites
- Turinia pagei8 sites
- Zascinaspis heintzi8 sites
- Nikolivia elongata7 sites
- Anglaspis6 sites
- Traquairaspis6 sites
- Homalaspidella5 sites
- Phialaspis symondsi5 sites
- Boreania minima4 sites
- Ctenaspis4 sites
- Machairaspis4 sites
- Nikolivia depressa4 sites
- Nostolepis4 sites
- Podolaspis lerichei4 sites
- Poraspis polaris4 sites
- Stegobranchiaspis baringensis4 sites
- Baringaspis dineleyi3 sites
- Canonia grossi3 sites
- Ctenaspis dentata3 sites
- Dinaspidella3 sites
The Paleobiology Database records 56 sites for Protopteraspididae, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
14 references- Protopteraspis Gosseleti (Vertebrata: Pteraspidomorphi: Heterostraci) from the Lower Devonian of Shropshire, England17
- The vertebrate stratigraphy of the Lower Devonian (Red Bay Group and Wood Bay Formation) of Spitsbergen12
- The Early Devonian armoured agnathans of Podolia, Ukraine8
- Upper Silurian and Devonian heterostracan pteraspidomorphs (Vertebrata) from Severnaya Zemlya (Russia): a preliminary report with biogeographical and biostratigraphical implications6
- New Pteraspididae (Agnatha, Heterostraci) from the Lower Devonian of Northwest Territories, Canada5
- Gigantaspis – a new genus of fam. Pteraspididae from Spitsbergen5
- Habitat of Llandoverian-Lochkovian eurypterids4
- Ostracoderm faunas of the Delorne and associated Siluri-Devonian Formations, North Western Territories, Canada2
- Chapter 4. Early Devonian fossil fishes sites of the Welsh Borders1
- The Furcacaudiformes: a new order of jawless vertebrates with thelodont scales, based on articulated Silurian and Devonian fossils from northern Canada1
- The Downtonian and Devonian vertebrates of Spitsbergen. XV* New Heterostracans from the Lower Devonian Red Bay Group, northern Spitsbergen1
- Zascinaspis laticephala nov. sp. (Agnatha, Heterostraci) du De´vonien infe´rieur du Spitsberg1
Each of the 65 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 2 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Protopteraspididae
- Source
- Wikipedia/Commons
- Credit
- Apokryltaros
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan of Protopteraspididae was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itPteraspidiformes through timeFormations of the DevonianThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Novitskaya 1983